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Glossary of leaf morphology





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The following terms are used to describe leaf morphology in the description and taxonomy of plants. Leaves may be simple (a single leaf blade or lamina) or compound (with several leaflets). The edge of the leaf may be regular or irregular, may be smooth or bearing hair, bristles or spines. For more terms describing other aspects of leaves besides their overall morphology see the leaf article.

Chart illustrating leaf morphology terms

The terms listed here all are supported by technical and professional usage, but they cannot be represented as mandatory or undebatable; readers must use their judgement. Authors often use terms arbitrarily, or coin them to taste, possibly in ignorance of established terms, and it is not always clear whether because of ignorance, or personal preference, or because usages change with time or context, or because of variation between specimens, even specimens from the same plant. For example, whether to call leaves on the same tree "acuminate", "lanceolate", or "linear" could depend on individual judgement, or which part of the tree one collected them from. The same cautions might apply to "caudate", "cuspidate", and "mucronate", or to "crenate", "dentate", and "serrate".

Another problem is to establish definitions that meet all cases or satisfy all authorities and readers. For example, it seems altogether reasonable to define a mucro as "a small sharp point as a continuation of the midrib", but it may not be clear how small is small enough, how sharp is sharp enough, how hard the point must be, and what to call the point when one cannot tell whether the leaf has a midrib at all. Various authors or field workers might come to incompatible conclusions, or might try to compromise by qualifying terms so vaguely that a description of a particular plant practically loses its value.

Use of these terms is not restricted to leaves, but may be applied to morphology of other parts of plants, e.g. bracts, bracteoles, stipules, sepals, petals, carpelsorscales. Some of these terms are also used for similar-looking anatomical features on animals.

Leaf structure

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Leaves of most plants include a flat structure called the blade or lamina, but not all leaves are flat, some are cylindrical. Leaves may be simple, with a single leaf blade, or compound, with several leaflets. In flowering plants, as well as the blade of the leaf, there may be a petiole and stipules; compound leaves may have a rachis supporting the leaflets. Leaf structure is described by several terms that include:[citation needed]

 
Bipinnate leaf anatomy with labels showing alternative usages
 
A ternate compound leaf with a petiole but no rachis (or rachillae)

Image

Term

Latin

Description

 

bifoliolate

Having two leaflets[1]

geminate

jugate

 

bigeminate

Having two leaflets, each leaflet being bifoliolate

 

bipinnate

bipinnatus

The leaflets are themselves pinnately-compound; twice pinnate

 

biternate

With three components, each with three leaflets

 

imparipinnate

With an odd number of leaflets, pinnate with a terminal leaflet (the opposite of paripinnate)

odd-pinnate

 

paripinnate

Pinnate with an even number of leaflets, lacking a terminal leaflet (the opposite of imparipinnate)

even-pinnate

 

palmately compound

palmatus

Consisting of leaflets all radiating from one point

pinnately compound

pinnatus

Having two rows of leaflets on opposite sides of a central axis, see imparipinnate and paripinnate

 

simple

Leaf blade in one continuous section, without leaflets (not compound)

 

ternate

ternatus

With three leaflets

trifoliate

trifoliatus

trifoliolate

trifoliolatus

 

tripinnate

tripinnatus

Pinnately compound in which each leaflet is itself bipinnate

Leaf and leaflet shapes

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Being one of the more visible features, leaf shape is commonly used for plant identification. Similar terms are used for other plant parts, such as petals, tepals, and bracts.

 
Oddly pinnate, pinnatifid leaves (Coriandrum sativum, coriander or cilantro)
 
Partial chlorosis revealing palmate venation in simple leaves of Hibiscus mutabilis

Image

Term

Latin

Refers principally to

Description

 

acicular

acicularis

whole leaf

Slender and pointed, needle-like

 

acuminate

acuminatus

leaf tip

Tapering to a long point in a concave manner

 

acute

leaf tip or base

Pointed, having a short sharp apex angled less than 90°

 

apiculate

apiculatus

leaf tip

Tapering and ending in a short, slender point

 

aristate

aristatus

leaf tip

Ending in a stiff, bristle-like point

asymmetrical

whole leaf

With the blade shape different on each side of the midrib

 

attenuate

attenuatus

leaf base

Having leaf tissue taper down the petiole to a narrow base and always having some leaf material on each side of the petiole

 

auriculate

auriculatus

leaf base

Having ear-shaped appendages reaching beyond the attachment to the petiole or stem (in case of a seated leaf)

 

caudate

caudatus

leaf tip

Tailed at the apex

cirrus, cirrate

leaf tip

Having a rachis that extends beyond the leaf blade or leaflets into a long whip-like extension or cirrus (common in climbing palms); antonym: ecirrate

 

cordate, cordiform

cordatus

whole leaf or base

Heart-shaped, with the petiole or stem attached to the notch

 

cuneate

cuneatus

leaf base

Triangular, wedge-shaped, stem attaches to point

cuneiform

whole leaf

Narrowly triangular, widest on the opposite end from the stem, with the corners at that end rounded

 

cuspidate

cuspidatus

leaf tip

With a sharp, elongated, rigid tip; tipped with a cusp

 

deltoid, deltate

deltoideus

whole leaf

Shaped like the Greek letter delta; triangular with stem attached to side

 

digitate

digitatus

whole leaf

A palmately compound leaf with leaflets, similar to palmate[2]

ecirrate

leaf tip

Without a cirrus; antonym: cirrate

 

elliptic

ellipticus

whole leaf

Shaped like an ellipse (widest at mid-blade and with similar convex tapering towards apex and base), with a short or no point

 

emarginate

emarginatus

leaf tip

Slightly indented at the tip

 

ensiform

ensiformis

whole leaf

Shaped like a sword; long and narrow with a sharp pointed tip

 

falcate

falcatus

whole leaf

Sickle-shaped

 

fenestrate

fenestratus

leaf surface features

Large openings through the leaf; see perforate; sometimes used to describe leaf epidermal windows

 

filiform

filiformis

whole leaf

Thread- or filament-shaped

 

flabellate

flabellatus

whole leaf

Semi-circular or fan-like

 

hastate

hastatus

whole leaf or base

Spear-shaped: pointed, with barbs, shaped like a spear point, with flaring pointed lobes at the base

 

laciniate

lacinatus

whole leaf

Very deeply lobed with the lobes being very drawn out and often making the leaf look somewhat like a branch or a pitchfork

laminar

3-D shape

Flat (like most leaves)

 

lanceolate

lanceolatus

whole leaf

Long, wider in the middle, shaped like a lance tip

 

linear

linearis

whole leaf

Long and very narrow like a blade of grass

 

lobed

lobatus

whole leaf

Being divided by clefts; may be pinnately lobedorpalmately lobed

 

lorate

loratus

whole leaf

Having the form of a thong or strap

 

lyrate

lyratus

whole leaf

Shaped like a lyre, pinnately lobed leaf with an enlarged terminal lobe and smaller lateral lobes. See also List of lyrate plants.

 

mucronate

mucronatus

leaf tip

Ending abruptly in a small sharp point as a continuation of the midrib[3]

 

multifid

multi + findere

whole leaf

Cleft into many parts or lobes

 

obcordate

obcordatus

whole leaf

Heart-shaped, stem attaches at the tapering end

 

oblanceolate

oblanceolatus

whole leaf

Much longer than wide and with the widest portion near the tip; reversed lanceolate

 

oblique

leaf base

Asymmetrical leaf base, with one side lower than the other

 

oblong

oblongus

whole leaf

Having an elongated form with slightly parallel sides; roughly rectangular

 

obovate

obovatus

whole leaf

Teardrop-shaped, stem attaches to the tapering end; reversed ovate

obtrullate

whole leaf

Reversed trullate; the longer sides meet at the base rather than the apex.

 

obtuse

obtusus

leaf tip or base

Blunt, forming an angle > 90°

 

orbicular

orbicularis

whole leaf

Circular

 

ovate

ovatus

whole leaf

Egg-shaped, with a tapering point and the widest portion near the petiole

 

palmate

palmatus

whole leaf

Palm-shaped, i.e. with lobes or leaflets stemming from the leaf base[4]

 

palmately lobed

palmatus

whole leaf

Lobes spread radially from a point[5]

 

palmatifid

palma + findere

whole leaf

Palm-shaped, having lobes with incisions that extend less than halfway toward the petiole

 

palmatipartite

palma + partiri

whole leaf

Having palmate lobes with incisions that extend over halfway toward the petiole

 

palmatisect

palma + secare

whole leaf

Having palmate lobes with incisions that extend almost up, but not quite to the petiole.

 

pandurate

panduratus

whole leaf

Fiddle-shaped; obovate with a constriction near the middle.

 

pedate

pedatus

whole leaf

Palmate, with cleft lobes[6]

 

peltate

peltatus

stem attachment

A round leaf where the petiole attaches near the center, e.g. a lotus leaf

 

perfoliate

perfoliatus

stem attachment

With the leaf blade surrounding the stem such that the stem appears to pass through the leaf

 

perforate

perforatus

leaf surface features

Many holes, or perforations, on leaf surface. Compare with fenestrate.

 

pinnately lobed

pinna + lobus

whole leaf

Having lobes pinnately arranged on the central axis

 

pinnatifid

pinna + findere

whole leaf

Having lobes with incisions that extend less than halfway to the midrib

 

pinnatipartite

pinnatus + partiri

whole leaf

Having lobes with incisions that extend more than halfway to the midrib

 

pinnatisect

pinnatus + sectus

whole leaf

Having lobes with incisions that extend almost to, or up to, the midrib

 

plicate

plicatus

3-D shape

Folded into pleats, usually lengthwise, serving the function of stiffening a large leaf

 

reniform

reniformis

whole leaf

Shaped like a kidney, with an inward curve on one side

 

retuse

leaf tip

With a shallow notch in a round apex

 

rhomboid, rhombic

rhomboidalis

whole leaf

Diamond-shaped

 

rounded

rotundifolius

leaf tip or base

Circular, no distinct point

semiterete

3-D shape

Rounded on one side and flat on the other

 

sinuate

sinuatus

3-D shape

Circularly-lobed leaves

 

sagittate

sagittatus

whole leaf

Arrowhead-shaped with the lower lobes folded, or curled downward

 

spatulate

spathulatus

whole leaf

Spoon-shaped; having a broad flat end which tapers to the base

 

spear-shaped

hastatus

whole leaf

See hastate.

 

subobtuse

subobtusus

leaf tip or base

Somewhat blunted; neither blunt nor sharp

 

subulate

subulatus

leaf tip

Awl-shaped with a tapering point

 

terete

3-D shape

Cylindrical with a circular or distorted circular cross-section and a single surface wrapping around it with no grooves or ridges. Subterete means the leaves are not completely terete, as seen in various lichens and succulents.

 

trullate

whole leaf

Shaped like a masonry trowel

 

truncate

truncatus

leaf tip or base

With a squared-off end

undulate

undulatus

3-D shape

Wave-like

 

unifoliate

unifoliatus

compound leaves

With a single leaflet; it is distinct from a simple leaf by the presence of two abscission layers and often by petiolules and stipels.

Edge

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Leaf margins (edges) are frequently used in visual plant identification because they are usually consistent within a species or group of species, and are an easy characteristic to observe. Edge and margin are interchangeable in the sense that they both refer to the outside perimeter of a leaf.

Image

Term

Latin

Description

 

entire

Forma
integra

Even; with a smooth margin; without toothing

 

ciliate

ciliatus

Fringed with hairs

 

crenate

crenatus

Wavy-toothed; dentate with rounded teeth

crenulate

crenulatus

Finely crenate

crisped

crispus

Curly

 

dentate

dentatus

Toothed;

may be coarsely dentate, having large teeth

orglandular dentate, having teeth which bear glands

 

denticulate

denticulatus

Finely toothed

 

doubly serrate

duplicato-dentatus

Each tooth bearing smaller teeth

 

serrate

serratus

Saw-toothed; with asymmetrical teeth pointing forward

 

serrulate

serrulatus

Finely serrate

 

sinuate

sinuosus

With deep, wave-like indentations; coarsely crenate

 

lobate

lobatus

Indented, with the indentations not reaching the center

 

undulate

undulatus

With a wavy edge, shallower than sinuate

 

spiny or pungent

spiculatus

With stiff, sharp points such as thistles

Leaf folding

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Leaves may also be folded, sculpted or rolled in various ways. If the leaves are initially folded in the bud, but later unrolls it is called vernation, ptyxis is the folding of an individual leaf in a bud.

Image

Term

Latin

Description

carinate or keeled

carinatus

With a longitudinal ridge, keel-shaped

 

conduplicate

Folded upwards, with the surfaces close to parallel

 

cucullate

Forming a hood, margins and tip curved downward

 

involute

Rolled upwards (towards the adaxial surface)

 

plicate

plicatus

With parallel folds

reduplicate

Folded downwards, with the surfaces close to parallel

 

revolute

Rolled downwards (towards the abaxial surface)

supervolute

Opposing left and right halves of lamina folded along longitudinal axis, with one half rolled completely within the other

Latin descriptions

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The Latin word for 'leaf', folium, is neuter. In descriptions of a single leaf, the neuter singular ending of the adjective is used, e.g. folium lanceolatum 'lanceolate leaf', folium lineare 'linear leaf'. In descriptions of multiple leaves, the neuter plural is used, e.g. folia linearia 'linear leaves'. Descriptions commonly refer to the plant using the ablative singular or plural, e.g. foliis ovatis 'with ovate leaves'.[7]

See also

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References

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  1. ^ Radford, A.E.; Dickison, W.C.; Massey, J.R.; Bell, C.R. (1976). "Phytography - Morphological Evidence". Vascular Plant Systematics. Harper and Row, New York.
  • ^ Index of Garden Plants, Mark Griffiths, Timber Press, 1992
  • ^ Mucronate Archived 2016-03-03 at the Wayback Machine, Answers.com, from Roget's Thesaurus.
  • ^ "palmate (adj. palmately)". GardenWeb Glossary of Botanical Terms. iVillage GardenWeb. 2006. Archived from the original on 13 February 2009. Retrieved 19 October 2008.
  • ^ Nelson, Randal C. (2009) [2012]. "Leaf description glossary". University of Rochester. Archived from the original on 1 August 2020. Retrieved 2 November 2017.
  • ^ "Pedate leaf". Retrieved 24 February 2014.
  • ^ Stearn (2004), pp. 439–440.
  • Bibliography

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  • "Leaves"., in Massey & Murphy (1996)
  • "Shapes"., in Massey & Murphy (1996)
  • Massey, Jimmy R.; Murphy, James C. (1996). "Vascular plant systematics". NC Botnet. University of North Carolina at Chapel Hill. Retrieved 19 January 2016.
  • edit

    Retrieved from "https://en.wikipedia.org/w/index.php?title=Glossary_of_leaf_morphology&oldid=1211276564#Leaf_and_leaflet_shapes"
     



    Last edited on 1 March 2024, at 19:07  





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