Raspberries, blackberries, and dewberries are common, widely distributed members of the genus, and bristleberries are endemic to North America. Most of these plants have woody stems with prickles like roses; spines, bristles, and gland-tipped hairs are also common in the genus. The Rubusfruit, sometimes called a bramble fruit, is an aggregateofdrupelets. The term "cane fruit" or "cane berry" applies to any Rubus species or hybrid which is commonly grown with supports such as wires or canes, including raspberries, blackberries, and hybrids such as loganberry, boysenberry, marionberry and tayberry.[6] The stems of such plants are also referred to as canes.
Bramble bushes typically grow as shrubs (though a few are herbaceous), with their stems being typically covered in sharp prickles.[7] They grow long, arching shoots that readily root upon contact with soil,[8] and form a soil rootstock from which new shoots grow in the spring.[9] The leaves are either evergreenordeciduous, and simple, lobed, or compound.[7] The shoots typically do not flower or set fruit until the second year of growth (i.e. they are biennial).[9] The rootstock is perennial.[10] Most species are hermaphrodites with male and female parts being present on the same flower.[7] Bramble fruits are aggregate fruits formed from smaller units called drupelets.[9]
Around 60-70% of species of Rubus are polyploid (having more than two copies of each chromosome), with species ranging in ploidy from diploid (2x, with 14 chromosomes[11]) to tetradecaploid (14x).[12]
Some treatments have recognized dozens of species each for what other, comparably qualified botanists have considered single, more variable species. On the other hand, species in the other Rubus subgenera (such as the raspberries) are generally distinct, or else involved in more routine one-or-a-few taxonomic debates, such as whether the European and American red raspberries are better treated as one species or two (in this case, the two-species view is followed here, with R. idaeus and R. strigosus both recognized; if these species are combined, then the older name R. idaeus has priority for the broader species).
The classification presented below recognizes 13 subgenera within Rubus, with the largest subgenus (Rubus) in turn divided into 12 sections. Representative examples are presented, but many more species are not mentioned here. A comprehensive 2019 study found subgenera Orobatus and Anoplobatus to be monophyletic, while all other subgenera to be paraphyleticorpolyphyletic.[13]
The genus has a likely North American origin,[13] with fossils known from the Eocene-aged Florissant Formation of Colorado, around 34 million years old.[14]Rubus expanded into Eurasia, South America, and Oceania during the Miocene.[13]Fossil seeds from the early MioceneofRubus have been found in the Czech part of the Zittau Basin.[15] Many fossil fruits of †Rubus laticostatus, †Rubus microspermus and †Rubus semirotundatus have been extracted from bore hole samples of the Middle Miocene fresh water deposits in Nowy Sacz Basin, West Carpathians, Poland.[16]
Molecular data have backed up classifications based on geography and chromosome number, but following morphological data, such as the structure of the leaves and stems, do not appear to produce a phylogenetic classification.[17]
The term "hybrid berry" is often used collectively for those fruits in the genus Rubus which have been developed mainly in the U.S. and U.K. in the last 130 years. As Rubus species readily interbreed and are apomicts (able to set seed without fertilisation), the parentage of these plants is often highly complex, but is generally agreed to include cultivars of blackberries (R. ursinus, R. fruticosus) and raspberries (R. idaeus). The British National Collection of Rubus stands at over 200 species and, although not within the scope of the National Collection, also hold many cultivars.[18][19]
The generic name means blackberry in Latin and was derived from the word ruber, meaning "red".[21]
The blackberries, as well as various other Rubus species with mounding or rambling growth habits, are often called brambles. However, this name is not used for those like the raspberry that grow as upright canes, or for trailing or prostrate species, such as most dewberries, or various low-growing boreal, arctic, or alpine species. The scientific study of brambles is known as "batology".
"Bramble" comes from Old English bræmbel, a variant of bræmel.[5]
^ ab"Rubus L."Plants of the World Online. Board of Trustees of the Royal Botanic Gardens, Kew. 2021. Archived from the original on 31 March 2023. Retrieved 4 September 2021.
^Acta Palaeobotanica – 43(1): 9-49, January 2003 – Early Miocene carpological material from the Czech part of the Zittau Basin – Vasilis Teodoridis
^Łańcucka-Środoniowa M.: Macroscopic plant remains from the freshwater Miocene of the Nowy Sącz Basin (West Carpathians, Poland) [Szczątki makroskopowe roślin z miocenu słodkowodnego Kotliny Sądeckiej (Karpaty Zachodnie, Polska)]. Acta Palaeobotanica 1979 20 (1): 3-117.
^Lawrence A. Alice & Christopher S. Campbell (1999). "Phylogeny of Rubus (rosaceae) based on nuclear ribosomal DNA internal transcribed spacer region sequences". American Journal of Botany. 86 (1). Botanical Society of America: 81–97. doi:10.2307/2656957. JSTOR2656957. PMID21680348.
Data related to Rubus at Wikispecies
