Pinus peuce (Macedonian pine[3]orBalkan pine[4]) (Serbo-Croatian and Macedonian: молика, molika; Bulgarian: бяла мура, byala mura) is a species of pine native to the mountains of North Macedonia, Bulgaria, Albania, Montenegro, Kosovo, the extreme southwest of Serbia, and the extreme north of Greece,[5][6][7] growing typically at (600-) 1,000-2,200 (-2,300) m altitude. It often reaches the alpine tree line in this area. The mature size is up to 35–40 m height, and 1.5 m trunk diameter.[5][6] However, the height of the tree diminishes strongly near the upper tree line and may even obtain shrub sizes.[8]
It is a member of the white pine group, Pinus subgenus Strobus, and like all members of that group, the leaves ('needles') are in fascicles (bundles) of five, with a deciduous sheath. They are 6–11 cm long. Its pine cones are mostly 8–16 cm long, occasionally up to 20 cm long, green at first, becoming yellow-brown when mature, with broad, flat to downcurved scales. The 6–7 mm long seeds have a 2 cm wing and can be wind-dispersed, but are also very often dispersed by spotted nutcrackers.[5][6]
The species was first described in scientific literature by August Grisebach in 1844, based on field observations of montane pines on Baba Mountain, above Bitola.[2]
Macedonian pine is one of the most valuable conifer species in the Balkan Peninsula. Its durable wood is highly valued in construction, furniture production, wood-carving and cooperage. The tree is also exceptionally good at adapting to severe mountain climate conditions, which makes it a valuable species for afforestation on high terrain for protection against erosion. The local population use P. peuceresin to cure wounds, pectoral, skin and stomach diseases, varicose veins and other illnesses.[8]
Macedonian pine is also a popular ornamental tree in parks and large gardens, giving reliable steady though not fast growth on a wide range of sites. It is very tolerant of severe winter cold, hardy down to at least -45 °C, and also of wind exposure. It is locally naturalisedinPunkaharju in eastern Finland.[6]
Like other European and Asian white pines, Macedonian pine is very resistant to white pine blister rust (Cronartium ribicola).[9][10] This fungal disease was accidentally introduced from Europe into North America, where it has caused severe mortality in the American native white pines (e.g. western white pine, sugar pine, whitebark pine) in many areas. Macedonian pine is of great value for research into hybridisation and genetic modification to develop rust resistance in these species; hybrids with eastern white pine inherit some resistance.[10]
Synonyms include Pinus cembra var. fruticosa Griseb.,[11]Pinus excelsa var. peuce (Griseb.) Beissn.,[12]Pinus peuce var. vermiculata Christ,[10] and Balkan pine.[13]
^Critchfield, W. B., & Little, E. L. (1966). Geographic distribution of the pines of the World. U.S. Dept. of Agriculture Forest Service Misc. Publ. 991.
^Søegaard, B. (1972). Relative blister rust resistance of native and introduced white pines in Europe. Pages 233–239 in Bingham, R. T. et al., eds. Biology of Rust Resistance in Forest Trees. U.S. Dept. of Agriculture Forest Service Misc. Publ. 1221.
^ abcPopnikola, N., M. Jovancevic, and M. Vidakovic. 1978. Genetics of Pinus peuce Gris. Annales Forestales 7/6: 187-206.
