Ciconia

Ciconia Temporal range: Early Miocene to present
Fledgling (left) and adult European white stork (Ciconia ciconia ciconia)
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Aves
Order:	Ciconiiformes
Family:	Ciconiidae
Genus:	Ciconia Brisson, 1760
Type species
Ardea ciconia Linnaeus, 1758
Species
Ciconia abdimii Ciconia boyciana Ciconia ciconia Ciconia episcopus Ciconia maguari Ciconia microscelis Ciconia nigra Ciconia stormi
Synonyms
Dissoura Euxenura Sphenorhynchus

Ciconia (/sɪˈkoʊ.ni.ə/ sih-KOH-nee-uh; Classical Latin: [kiˈkoː.ni.a]) is a genus of birds in the stork family. Six of the seven living species occur in the Old World, but the maguari stork has a South American range. In addition, fossils suggest that Ciconia storks were somewhat more common in the tropical Americas in prehistoric times.

The genus was introduced by the French zoologist Mathurin Jacques Brisson in 1760 with the white stork (Ciconia ciconia) as the type species.^[1][2] The genus name is the Latin word for "stork",^[3] and was originally recorded in the works of Horace and Ovid.^[4]

The Abdim's stork is the smallest of the family, but other species in the genus are generally medium-sized storks, with long legs and a long thick bill. The members of this genus are more variable in plumage than other stork genera, but all species are black (at least to the wings) and white (at least underparts or neck). Juveniles are a duller, browner version of the adult.

Depending on species, breeding can be in solitary pairs or colonies. Pairs usually stay together for life. They typically build large stick nests in trees, although the Abdim's stork sometimes will nest on cliffs, the maguari stork will nest on the ground and at least three species will construct their nests on human habitations. One of these, the white stork, is probably the best known of all storks, with a wealth of legend and folklore associated with this familiar summer visitor to Europe.

These storks feed on frogs, insects, fish, crustaceans, small birds, lizards and rodents. They fly with the neck outstretched, like most other storks, but unlike herons which retract their neck in flight.

The migratory species like the white stork and the black stork soar on broad wings and rely on thermals of hot air for sustained long distance flight. Since thermals only form over land, these storks, like large raptors, must cross the Mediterranean at the narrowest points, and many of these birds can be seen going through the Straits of Gibraltar and the Bosphorus on migration.

The genus contains eight extant species:^[5]

Image	Scientific name	Common Name	Distribution
	Ciconia abdimii	Abdim's stork	Widespread in open habitats of Sub-Saharan Africa, and in Yemen. Breeds in northern half of range and spends non-breeding period in southern half
	Ciconia episcopus	Asian woolly-necked stork	Southern Asia, from Pakistan to Indonesia and the Philippines
	Ciconia stormi	Storm's stork	Borneo, Sumatra and the Thai-Malay Peninsula
	Ciconia nigra	Black stork	Breeds from Eastern Asia (Siberia and northern China) west to Central and Southern Europe. Winters in South, Southeast and East Asia, and in tropical Africa. A resident (non-migratory) population in southern Africa
	Ciconia maguari	Maguari stork	Widespread in open wetland habitats in northern, central and southern South America
	Ciconia microscelis	African woolly-necked stork	tropical Africa
	Ciconia ciconia	White stork	Breeds in Europe to central Asia, and in northern Africa. Winters in Sub-Saharan Africa and South Asia
	Ciconia boyciana	Oriental stork	Breeds in Russian Far East and northeast China. Winters in Japan, Korean Peninsula, east-central and southeast China, and Taiwan

The fossil record of the genus is extensive, indicating that Ciconia storks were once more widespread than they are today. Although the known material tends to suggest that the genus evolved around the Atlantic, possibly in western Europe or Africa, the comparative lack of fossil sites in Asia makes this assumption not well-founded presently. All that can be said is that by the Early Pliocene, Ciconia was widespread at least all over the Northern Hemisphere.

Fossil members of the genus include:

• Ciconia louisebolesae (Early Miocene of Riversleigh, Australia)
• ?Ciconia minor (Early Miocene of Rusinga Island, Kenya)
• ?Ciconia sarmatica (Late Miocene of Credinţa, Romania)
• ?Ciconia gaudryi (Late Miocene/Early Pliocene of Pikermi, Greece)
• Ciconia sp. 1 (Late Miocene/Early Pliocene of Lee Creek Mine, the United States)
• Ciconia sp. 2 (Late Miocene/Early Pliocene of Lee Creek Mine, the United States)
• ?Ciconia kahli (Early Pliocene of South Africa)
• Ciconia lucida (Mongolian stork), (Middle Pliocene of Mongolia)
• Ciconia maltha (asphalt stork or La Brea stork), (Late Pliocene – Late Pleistocene of the western and southern United States, Cuba and Bolivia)
• Ciconia stehlini (Late Pliocene – Early Pleistocene of Hungary) – may belong to extant species
• Ciconia nana (Australian stork) – (Early to Middle Pliocene, Late Pleistocene of Australia) – formerly Xenorhynchus^[6]
• Ciconia sp. (Late Pleistocene/Early Holocene of Las Breas de San Felipe, Cuba)^[7]

Adistal radiusinLate Pleistocene deposits of San Josecito Cavern (Mexico) may belong in this genus or in Mycteria; it is smaller than that of any known American stork, Ciconia or otherwise.^[8] The proposed fossil genus Prociconia from Brazil, also of Late Pleistocene age, may be a junior synonym of either this genus or Jabiru. A distal tarsometatarsus found in a rock shelteronRéunion was probably of a bird taken there as food by early settlers; no known account mentions the presence of storks on the Mascarenes, and while this subfossil was initially believed to be from a stork, it is today assigned to the Réunion ibis (Threskiornis solitarius) which is quite similar to storks osteologically and was not yet described when the bone was discovered (Cowles, 1994).

• Barlow, Clive (1997): A field guide to birds of the Gambia and Senegal. Pica Press, Nr. Robertsbridge (East Sussex). ISBN 1-873403-32-1
• Cowles, Graham S. (1994): A new genus, three new species and two new records of extinct Holocene birds from Réunion Island, Indian Ocean. Geobios 27(1): 87–93.
• Grimmett, Richard; Inskipp, Carol, Inskipp, Tim & Byers, Clive (1999): Birds of India, Pakistan, Nepal, Bangladesh, Bhutan, Sri Lanka, and the Maldives. Princeton University Press, Princeton, N.J.. ISBN 0-691-04910-6
• Hilty, Steven L. (2003): Birds of Venezuela. Christopher Helm, London. ISBN 0-7136-6418-5

Wikispecies has information related to Ciconia.

Wikimedia Commons has media related to Ciconia.

• ARKive: images and movies of Ciconia boyciana
• Ciconia videos on Internet Bird Collection:
• Ecological education through live white storks' nest webcam observation Archived 2014-07-14 at the Wayback Machine