Najas marina is a species of aquatic plant known by the common names spiny water nymph, spiny naiad and holly-leaved naiad. It is an extremely widespread species, reported across Europe, Asia, Africa, Australia, the Americas and many oceanic islands. It can be found in many types of freshwater and brackish aquatic habitat, including bodies of alkaline water.[3][4][5]
Najas marina is an annual producing a slender, branching stem up to 40 or 45 centimeters in maximum length. The evenly spaced leaves are up to 4 centimeters long, 1 to 3 millimeters wide, and edged in tiny sawlike teeth. The leaf has prickles along its midvein. Minute stalkless, green flowers occur in the leaf axils. The plant is dioecious, with male and female flower types occurring on separate individuals.[6][4][5][7][8] In the British Isles it is possible that only female plants occur. It flowers in mid-summer.[9]
Najas marina has a wide, almost circumglobal distribution in temperate and tropical regions. It occurs in mesoeutrophic water over deep peat or mud. It was first recorded in the British Isles in 1883 at Hickling Broad in Norfolk where it had become established. Populations declined in the 1960s because of pollution, but action has been taken to reduce the level of nutrients in the Norfolk Broads and the water quality has improved.[11]
^"Najus marina". Online Atlas of the British and Irish Flora. Archived from the original on 30 September 2021. Retrieved 14 March 2020.
^Łańcucka-Środoniowa M.: Macroscopic plant remains from the freshwater Miocene of the Nowy Sącz Basin (West Carpathians, Poland) [Szczątki makroskopowe roślin z miocenu słodkowodnego Kotliny Sądeckiej (Karpaty Zachodnie, Polska)]. Acta Palaeobotanica 1979 20 (1): 3-117.
