Pan-Euungulata (literally: "all true ungulates"; also known as: Euungulata [sensu lato][2]) is a cladeofplacental mammals from grandorder Ferungulata, consisting of the taxon Euungulata [sensu stricto] and all taxa (species) more closely related to it than to any other living species.[1]
The specification of the above-mentioned taxa related to Euungulata s.s. varies from author to author and changes over time.[1][3][4][5] The extinct family Protungulatidae is a typical representative of such taxa.[2][6]
Some (usually older) texts use the conceptually similar clade Ungulata in the broadest sense (and/or the clade Ungulatomorpha, which is Ungulata in the broadest sense plus Zhelestidae). This clade differs from Pan-Euungulata in that it also includes the taxon Paenungulata (i.e. for example Proboscidea, Sirenia and Hyracoidea) and sometimes also the taxa Tubulidentata and Dinocerata (and in the case of Ungulatomorpha also the taxon Zhelestidae).[7][8][9]
Speculative reconstructionofProtungulatum, a possible ungulate relative
^ abcO’Leary, M. A., Bloch JI, Flynn, J. J., Gaudin, T. J., Giallombardo, A., Giannini, N. P., Goldber, S. L, Kraatz, B. P., Luo, Z-X, Jin Meng, Xijun Ni, Novacek, M. J., Perini, F. A., Randall, Z. S., Rougier, G. W., Sargis, E. J., Silcox, M. T., Simmons, N. B., Spaulding, M. Velazco, P. M., Weksler, M., Wible, J. R. Cirranello, A. L. (2013.) "The Placental Mammal Ancestor and the Post–K-Pg Radiation of Placentals." Science 339:6120:662-667. (Including Supplementary Material)
^Michelle Spaulding, Maureen A. O'Leary, John Gatesy - Spaulding M, O'Leary MA, Gatesy J (2009) Relationships of Cetacea (Artiodactyla) Among Mammals: Increased Taxon Sampling Alters Interpretations of Key Fossils and Character Evolution. PLoS ONE 4(9): e7062. doi:10.1371/journal.pone.0007062
^Averianov, Alexander, Archibald, J. David. Mammals from the mid-Cretaceous Khodzhakul Formation, Kyzylkum Desert, Uzbekistan. Cretaceous Research 26 (2005). page 595
