Verbena hastata, commonly known as American vervain,[1]blue vervain,[2]simpler's joy,[3]orswamp verbena,[4] is a perennialflowering plant in the vervain family Verbenaceae. It grows throughout the continental United States and in much of southern Canada.
V. hastata grows as a stiffly erect stem, occasionally branching in the upper half, reaching up to 1.5 m (5 ft) tall. The stems are four-angled (square), hairy, and green to reddish in color. Leaves are opposite, simple, and measure up to 15 cm (6 in) long and 3 cm (1 in) across.[5] They have doubly-serrate margins and a variety of shapes, from lanceolatetoovate, and may have 2 lateral lobes.[6]
The inflorescence is a panicle, or group, of flowering spikes, up to 30 cm (1 ft) long at the end of the upper stems. Each flowering spike in the panicle is up to 13 cm (5 in) long, with densely packed, numerous 5-lobed flowers, which measure up to 1 cm (0.25 in) long.[5] The flowers are violet or deep purple, rarely white.[6] They open from the bottom of the spike upward, with only a ring of a few flowers open at a time.[7]
In the recent evolutionary past, there has been an incident of chloroplast transfer of one of the latter or the swamp verbena to the mock vervainGlandularia bipinnatifida which is a close relative of the genusVerbena. It is unknown by what mechanism this happened, but it is suspected that hybridization is not responsible.[9]
V. hastata is native in the United States in all states except Alaska and Hawaii. In Canada, it is native in the provinces of British Columbia, Saskatchewan, Manitoba, Ontario, Quebec, New Brunswick, Nova Scotia, and Prince Edward Island.[4] The plant prefers moist conditions and typically grows in wet meadows, wet river bottomlands, stream banks, slough peripheries, fields and waste areas.[7] It is adapted to wetlands and can spread to form small colonies.[11]
