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(Top) 1 Plants 1.1 Ferns and fern allies 1.2 Angiosperms 2 Arthropods 2.1 Crustaceans 2.2 Insects 3 Molluscs 3.1 Cephalopods 4 Tetrapodomorphs 5 Amphibians 5.1 Newly named temnospondylians 5.2 Newly named amphibians 6 Ichthyosaurs 7 Lepidosauromorphs 7.1 Newly named basal lepidosauromorphs 7.2 Newly named plesiosaurs 7.3 Newly named squamates 8 Turtles 9 Archosauromorphs 9.1 Newly named crurotarsans 9.2 Newly named dinosaurs 9.3 Newly named birds 9.4 Newly named pterosaurs 10 Synapsids 10.1 Non-mammalian 10.2 Mammals 11 Trace fossils 12 References

2006 in paleontology

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List of years in paleontology

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Paleontology or palaeontology is the study of prehistoric life formsonEarth through the examination of plant and animal fossils.^[1] This includes the study of body fossils, tracks (ichnites), burrows, cast-off parts, fossilised feces (coprolites), palynomorphs and chemical residues. Because humans have encountered fossils for millennia, paleontology has a long history both before and after becoming formalized as a science. This article records significant discoveries and events related to paleontology that occurred or were published in the year 2006.

Plants[edit]

Ferns and fern allies[edit]

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes
Dickwhitea^[2]	Gen et sp nov	Valid	Karafit et al.	Eocene Ypresian	Eocene Okanagan Highlands Allenby Formation Princeton Chert	Canada British Columbia	Anathyriaceous fern The type species is D. allenbyensis
Trawetsia^[3]	Gen et sp nov	Valid	Smith et al.	Eocene Ypresian	Eocene Okanagan Highlands Allenby Formation Princeton Chert	Canada British Columbia	Anathyriaceous fern The type species is T. princetonensis
Wessiea oroszii^[4]	Sp nov	Valid	Serbet & Rothwell	Campanian	Horseshoe Canyon Formation	Canada Alberta	Afern of uncertain affinity

Angiosperms[edit]

Name	Novelty	Status	Authors	Age	Type locality	Location	Notes	Images
Beardia^[5]	Gen et sp nov	valid	Elliot, Mindell, & Stockey	Eocene	Appian Way Flora	Canada British Columbia	Ajuglandaceous fruit.

Arthropods[edit]

Crustaceans[edit]

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Stenochirus vahldieki^[6]	Sp nov	Valid	Schweigert & Garassino	Early Jurassic Callovian	La Voulte-sur-Rhöne	France	Astenochirid.

Insects[edit]

Name	Novelty	Status	Authors	Age	Type unit	Location	Notes	Images
Avitomyrmex^[7]	Gen et 3 sp. nov	valid	Archibald, Cover, & Moreau	Eocene Ypresian	Eocene Okanagan Highlands Tranquille Formation	Canada British Columbia	Amyrmeciine bulldog ant The type species is A. elongatus Other species are A. mastax and A. systenus
Emiliana^[8]	Trib, Gen, et sp. nov	valid	Shcherbakov	Eocene Ypresian	Green River Formation Parachute Member	USA Colorado	Atropiduchid planthopper
Eocenomyrma^[9]	Gen et 3 sp nov	Valid	Dlussky & Radchenko	Eocene Priabonian	Baltic amber	Russia	AMyrmicin ant The type species is E. orthospina Other species are E. electrina & E. elegantula	Eocenomyrma elegantula
Feroseta^[10]	Gen et sp nov	valid	Poinar	Miocene Burdigalian	Dominican amber	Dominican Republic	Amantidfly The type species is F. prisca
Macabeemyrma^[7]	Gen et sp. nov	valid	Archibald, Cover, & Moreau	Eocene Ypresian	Eocene Okanagan Highlands Tranquille Formation	Canada British Columbia	Amyrmeciine bulldog ant The type species is M. ovata	Macabeemyrma ovata
Palaeopsychops douglasae^[11]	Sp nov	valid	Archibald & Makarkin	Eocene Ypresian	Eocene Okanagan Highlands Coldwater Beds	Canada British Columbia	Anithionid giant lacewing
Palaeopsychops marringerae^[11]	Sp nov	valid	Archibald & Makarkin	Eocene Ypresian	Eocene Okanagan Highlands Klondike Mountain Formation	United States Washington	Anithionid giant lacewing	Palaeopsychops marringerae
Palaeopsychops quadratus^[11]	Sp nov	valid	Archibald & Makarkin	Eocene Ypresian	Fur Formation	Denmark	Anithionid giant lacewing
Palaeopsychops setosus^[11]	Sp nov	valid	Archibald & Makarkin	Eocene Ypresian	Eocene Okanagan Highlands Horsefly Shales	Canada British Columbia	Anithionid giant lacewing
Palaeopsychops timmi^[11]	Sp nov	valid	Archibald & Makarkin	Eocene Ypresian	Eocene Okanagan Highlands Klondike Mountain Formation	United States Washington	Anithionid giant lacewing	Palaeopsychops timmi
Palaeovespa menatensis^[12]	Sp nov	valid	Nel & Auvray	Paleocene Thanetian	Menat Formation	France	Avespid wasp	Palaeovespa menatensis
Polystoechotites^[11]	Gen et 3 sp nov	valid	Archibald & Makarkin	Eocene Ypresian	Eocene Okanagan Highlands Klondike Mountain Formation	United States Washington	Anithionid giant lacewing parataxon Species include P. barksdalae, P. falcatus, P. lewisi Also includes Polystoechotes piperatus (1908)	Polystoechotites barksdalae
Ypresiomyrma^[7]^[13]	Gen, sp., et comb nov	valid	Archibald, Cover, & Moreau	Eocene Ypresian	Fur Formation	Denmark	Amyrmeciine bulldog ant The type species is Y. orbiculata Other species are Y. bartletti & Pachycondyla rebekkae (1999)	Ypresiomyrma rebekkae

Molluscs[edit]

Cephalopods[edit]

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Tirolonautilus feltgeni^[14]	Sp nov	Valid	Chirat, Vaslet & Le Nindre	Late Permian	Khuff Formation	Saudi Arabia	Anautiloid.

Tetrapodomorphs[edit]

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Tiktaalik^[15]	Gen et sp nov	Valid	Daeschler, Shubin, & Jenkins	Late Devonian	Fram Formation	Canada ( Nunavut)	Type species T. roseae. The Earliest Tetrapod.	Tiktaalik roseae

Amphibians[edit]

Newly named temnospondylians[edit]

Name	Status	Authors	Age	Unit	Location	Notes	Images
Iberospondylus^[16]	Valid	Laurin Soler-Gijon	Late Carboniferous		Spain	The type species is Iberospondylus schultzei.	Iberospondylus
Tirraturhinus^[17]	Valid	Nield Damiani Warren	Early Triassic	Arcadia Formation	Australia	The type species is Tirraturhinus smisseni.
Trematolestes^[18]	Valid	Schoch	Middle Triassic	Erfurt Formation	Germany	The type species is Trematolestes hagdorni.	Trematolestes

Newly named amphibians[edit]

Name	Status	Authors	Age	Unit	Location	Notes	Images
Pangerpeton^[19]	Valid	Wang Evans	Late Jurassic/Early Cretaceous	Daohugou Beds	China	The type species is Pangerpeton sinensis.

Ichthyosaurs[edit]

Name	Status	Authors	Age	Unit	Location	Notes	Images
Maiaspondylus^[20]	Valid	Maxwell Caldwell	Early Cretaceous	Loon River Formation	Canada	The type species is Maiaspondylus lindoei.

Lepidosauromorphs[edit]

Newly named basal lepidosauromorphs[edit]

Name	Status	Authors	Age	Unit	Location	Notes	Images
Rautiania^[21]	Valid	Bulanov Sennikov	Late Permian	Vyasovka Formation	Russia	Two species was descript: Rautiania alexandri (type) and R. minchi.

Newly named plesiosaurs[edit]

Name	Status	Authors	Age	Unit	Location	Notes
Bobosaurus^[22]	Valid	Dalla Vecchia	Late Triassic	Rio del Lago Formation	Italy	The type species is Bobosaurus forojuliensis.
Futabasaurus^[23]	Valid	Sato Hasegawa Manabe	Late Cretaceous	Tamayama Formation	Japan	The type species is Futabasaurus suzukii.
Opallionectes^[24]	Valid	Kear	Early Cretaceous		Australia	The type species is Opallionectes andamookaensis.
Umoonasaurus^[25]	Valid	Kear Schroeder Lee	Early Cretaceous	Bulldog Shale	Australia	The type species is Umoonasaurus demoscyllus.

Newly named squamates[edit]

Name	Novelty	Status	Authors	Age	Unit	Location	Notes	Images
Najash^[26]	Gen et sp nov	Valid	Apesteguia & Zaher	Late Cretaceous	Candeleros Formation	Argentina	Type sp N rionegrina.

Turtles[edit]

Name	Status	Authors	Age	Unit	Location	Notes	Images
Gigantatypus^[27]	Valid	Kaddumi	Late Cretaceous (Maastrichtian)	Muwaqqar Chalk Marl Formation	Jordan	Asea turtle. The type species is G. salahi.

Archosauromorphs[edit]

Newly named crurotarsans[edit]

Name	Status	Authors	Age	Unit	Location	Notes	Images
Adamantinasuchus^[28]	Valid	Nobre Carvalho	Late Cretaceous	Adamantina Formation	Brazil	Anotosuchian. The type species is Adamantinasuchus navae.	Adamantinasuchus
Aldabrachampsus^[29]	Valid	Brochu	Pleistocene		Seychelles	Acrocodylid. The type species is Aldabrachampsus dilophus.
Effigia^[30]	Valid	Nesbitt Norell	Late Triassic	Chinle Formation	USA ( New Mexico)	Arauisuchian. The type species is Effigia okeeffeae.	Effigia
Isisfordia^[31]	Valid	Salisbury Molnar Frey Willis	Early-Late Cretaceous	Winton Formation	Australia	Aeusuchian. The type species is Isisfordia duncani.	Isisfordia

Newly named dinosaurs[edit]

Vickaryous, M K., 2006, New information on the cranial anatomy of Edmontonia rugosidens Gilmore, a Late Cretaceous nodosaurid dinosaur from Dinosaur Provincial Park, Alberta: JVP, v. 26, n. 4: 1011–1013.

Data are courtesy of George Olshevky's dinosaur genera list.^[32]

Name	Status	Authors		Location	Notes	Images
Adamantisaurus^[33]	Valid taxon	Rodrigo Santucci	Reinaldo Jose Bertini	Brazil		Adamantisaurus
Agujaceratops	Valid taxon	Spencer G. Lucas Sullivan	Hunt	USA ( Texas)		Agujaceratops
Alaskacephale^[34]	Valid taxon	Sullivan		USA ( Alaska)	Apachycephalosaurid from Alaska.	Alaskacephale
Aniksosaurus^[35]	Valid taxon	Martínez	Novas	Argentina
Antarctopelta.^[36]	Valid taxon	Salgado	Gasparini	Antarctica	An Ankylosaur from Antarctica	Antarctopelta
Balochisaurus	Valid taxon	Malkani		Pakistan
"Bayosaurus"	Nomen nudum	Rodolfo Coria Phillip J. Currie	Paulina Carabajal
Cedrorestes^[37]	Valid taxon	David Gilpin Tony DiCroce	Kenneth Carpenter	USA( Utah)
Dracorex^[38]	Jr. Synonym	Robert Bakker Sullivan Porter	Larson Saulsbury		Jr. Synonym of Pachycephalosaurus.	Dracorex
Dracovenator^[39]	Valid taxon	Yates		South Africa	An African Neotheropod related to Dilophosaurus.	Dracovenator
Erketu^[40]	Valid taxon	Ksepka	Mark Norell	Mongolia
Europasaurus^[41]	Valid taxon	Mateus Laven Knötschke vide: Sander Mateus	Laven Knötschke	Germany	A basal macronarian sauropod exhibiting insular dwarfism.	Europasaurus
Fusuisaurus^[42]	Valid taxon	Mo Wang W. Huang Z.	Huang X. Xu X.	China
Guanlong^[43]	Valid taxon	Xu Xing Clark Forster Mark Norell	Erickson Eberth Jia C. Zhao Q.	China	A basal, crested Tyrannosauroid.	Guanlong
Huanghetitan.^[44]	Valid taxon	You Li D.	Zhou L. Ji Q.	China
Jiutaisaurus^[45]	Valid taxon	Wu W. H. Dong Zhiming Sun	Li C. Li T.
Juravenator^[46]	Valid taxon	Göhlich	Luis M. Chiappe	Germany	A feathered Compsognathid.	Juravenator
Khetranisaurus^{[citation needed]}	Valid taxon	Malkani		Pakistan
Koutalisaurus^[47]	Jr. Synonym	Prieto-Marquez Gaete Rivas	Galobart Boada		Jr. Synonym of Pararhabdodon.
Ligabuesaurus^[48]	Valid taxon	Jose Bonaparte González-Riga (as González Riga)	Apesteguía	Argentina	An Argentine Titanosaur.
Mantellisaurus^[49]	Valid taxon	Paul		Belgium Germany UK Spain		Mantellisaurus
Mapusaurus^[50]	Valid taxon	Rodolfo Coria	Phillip J. Currie	Argentina	An Argentine Carcharodontosaur.	Mapusaurus
Marisaurus^{[citation needed]}	Valid taxon	Malkani		Pakistan
Maxakalisaurus^[51]	Valid taxon	Kellner Campos Azevedo Trotta	Henriques Craik Silva	Brazil		Maxakalisaurus
Othnielosaurus^[52]	Valid taxon	Peter Galton		USA ( Colorado, Utah and Wyoming)		Othnielosaurus
Pakisaurus^{[citation needed]}	Valid taxon	Malkani		Pakistan
Razanandrongobe^[53]	Valid taxon	Maganuco Dal Sasso	Pasini		Described by the authors as a possible crocodylomorph or theropod.
Sacisaurus^[54]	Valid taxon	Ferigolo	Langer	Brazil	Possibly a Dinosaur.	Sacisaurus
Sonidosaurus	Valid taxon	Xu X. Zhang X. H. Tan Q.	Zhao X. Tan L.	China
Sulaimanisaurus^{[citation needed]}	Valid taxon	Malkani		Pakistan
Theiophytalia^[55]	Valid taxon	Brill	Carpenter	USA ( Colorado)		Theiophytalia
Tsaagan^[56]	Valid taxon	Mark Norell Clark Turner	Makovicky Rinchen Barsbold Rowe	Mongolia		Tsaagan
Turiasaurus^[57]	Valid taxon	Royo-Torres Cobos	Alcalá	Spain
Vitakridrinda.^{[citation needed]}	Valid taxon	Malkani		Pakistan	An Abelisaur from Pakistan.
Xuanhuaceratops^[58]	Valid taxon	Zhao X. Cheng Z.	Xu X. Makovicky	China
Yamaceratops^[59]	Valid taxon	Makovicky	Mark Norell	China		Yamaceratops
Yinlong^[60]	Valid taxon	Xu X. Forster	Clark Mo	China		Yinlong
Yuanmousaurus^[61]	Valid taxon	Lü Li S. Ji Q.	Wang G. F. Zhang J. H. Dong Zhiming	China
Zapalasaurus	Valid taxon	Salgado Carvalho	Garrido	Argentina	An Argentine Rebbachisaur.

Newly named birds[edit]

Name	Status	Novelty	Authors	Age	Unit	Location	Notes
Abdounornis marinus ^[62]	Nomen Nudum	Thesis name	Bourdon	Early Ypresian		Morocco	A species of Charadriiformes, Abdounornithidae.
Aequornis traversei ^[62]	Nomen Nudum	Thesis name	Bourdon	Middle Eocene		Togo	A species of Pelagornithidae.
Anas cheuen ^[63]	Valid	Sp. nov.	Agnolín	Early or Middle Pleistocene	Miramar Formation	Argentina	A species of Anas.
Archaeorhynchus spathula ^[64]	Valid	Gen et Sp. nov.	Zhonghe & Fucheng	Early Cretaceous	Yixian Formation	China	AnOrnithurae, type sp A. spathula.
Branta thessaliensis ^[65]	Valid	Sp. nov.	Boev & Koufos	Middle Turolian		Greece	AnAnatidae.
Cerebavis cenomanica ^[66]	Valid	Gen. et Sp. nov.	Kurochkin, Saveliev, Postnov, Pervushov, & Popov	Cenomanian		Russia	AnEnantiornithes, described from a brain endocast.
Corvitalusoides ^[67]	Valid	Gen. et Sp. nov.	Boles	Late Oligocene or Early Miocene	Riversleigh	Australia: Queensland	APasseriformes Incertae Sedis. Type species C. grandiculus
Cyanoliseus patagonopsis ^[68]	Valid	Sp. nov.	Hospitaleche & Tambussi	Pleistocene	Miramar Formation	Argentina	APsittacidae.
Dalingheornis ^[69]	Valid	Gen. nov. et Sp. nov.	Zhang, Hou, Hasegawa, O'Connor, Martin, & Chiappe	Early Cretaceous	Yixian Formation	China	AnEnantiornithes, type species is Dal. liweii.
Dapingfangornis ^[70]	Valid	Gen. nov. et Sp. nov.	Li, Duan, Hu, Wang, Cheng, & Hou	Early Cretaceous	Jiufotang Formation	China	AnEnantiornithes, type species Dap. sentisorhinus
Didunculus placopedetes ^[71]	Valid	Sp. nov.	Steadman	Subrecent	'Eua	Tonga	AColumbidae.
Ducula harrisoni ^[72]	Valid	Sp. nov.	Wragg & Worthy	Subrecent	Henderson Island	Pitcairn	AColumbidae.
Eclectus infectus ^[73]	Valid	Sp. nov.	Steadman	Holocene	'Eua, 'Anatu	Vanuatu	APsittacidae.
Gallirallus ernstmayri ^[74]	Valid	Sp. nov.	Jeremy J. Kirchman & Steadman	Holocene	New Ireland, Bismarck Archipelago	Papua New Guinea	ARallidae.
Gallirallus pendiculentus ^[74]	Valid	Sp. nov.	Kirchman & Steadman	Holocene	Tinian	Northern Mariana Islands	ARallidae.
Gallirallus pisonii ^[74]	Valid	Sp. nov.	Kirchman & Steadman	Holocene	Tinian	Northern Mariana Islands	ARallidae.
Gallirallus storrsolsoni ^[75]	Valid	Sp. nov.	Kirchman & Steadman	Holocene	Huahine, Society Islands	French Polynesia	ARallidae.
Gallirallus temptatus ^[74]	Valid	Sp. nov.	Kirchman & Steadman	Holocene	Rota	Northern Mariana Islands	ARallidae.
Jungornis geraldmayri ^[76]	Valid	Sp. nov.	Mourer-Chauviré & Sigé	Late Eocene	Phosphorites du Quercy	France	AJungornithidae
Loxioides kikuchi ^[77]	Valid	Sp. nov.	James & Olson	Subrecent	Kaua'i, Hawaii Islands	USA	AFringillidae
Masillaraptor ^[78]	Valid	Gen. nov. et Sp. nov.	Mayr	Middle Eocene	Messel pit	Germany	AMasillaraptoridae, type species M. parvunguis
Nyctanassa carcinocatactes ^[79]	Valid	Sp. nov.	Olson & Wingate	Holocene	Bermuda	Bermuda	AnArdeidae.
Odontopteryx gigas ^[62]	Nomen Nudum	Thesis name	Bourdon	Early Ypresian		Morocco	AnOdontopterygidae
Odontoptila inexpectata ^[62]	Nomen Nudum	Thesis name	Bourdon	Early Ypresian		Morocco	AnOdontopterygidae
Pleistoanser ^[63]	Valid	Gen. et Sp. nov.	Agnolín	Early or Middle Pleistocene	Miramar Formation	Argentina	AnAnatidae. Type species is P. bravardi.
Plesiocathartes major ^[80]	Valid	Sp. nov.	Weidig	Late Wasatchian	Green River Formation	USA	ALeptosomidae.
Plesiocathartes wyomingensis ^[80]	Valid	Sp. nov.	Weidig	Late Wasatchian	Green River Formation	USA	ALeptosomidae.
Porphyrio mcnabi ^[75]	Valid	Sp. nov.	Kirchman & Steadman	Holocene	Huahine, Fa'ahia, Society Islands	French Polynesia	ARallidae.
Pygoscelis calderensis ^[81]	Valid	Sp. nov.	Hospitaleche, Chávez, & Fritis	Late Miocene-Middle Pliocene	Bahia Inglesa Formation	Chile	ASpheniscidae.
Pygoscelis grandis ^[82]	Valid	Sp. nov.	Walsh & Suárez	?Early Pliocene	Bahia Inglesa Formation	Chile	ASpheniscidae.
Shanxiornis ^[83]	Valid	Gen. et Sp. nov.	Wang, Zhao, Hu, & Sun	Early Pleistocene	Sanmen Formation	China	APhasianidae, type species is S. fenyinis
Tonniornis ^[84]	Valid	Gen. nov. et Sp. nov.	Tambussi, Hospitaleche, Reguero, & Marenssi	Late Eocene	La Meseta Formation	Antarctica	ASpheniscidae, two species T. mesetaensis & T. minimum
Waimanu ^[85]	Valid	Gen. nov. et Sp. nov.	Slack, Jones, Ando, Harrison, Fordyce, Arnason, & Penny	Early Paleocene	Basic Waipara Greensand	New Zealand	A stem Spheniscidae. The type species is Wa. manneringi. Originally the second species, Wa. tuatahi, was assigned to this genus as well; Mayr et al. (2018) transferred this species to the separate genus Muriwaimanu.^[86]
Wieslochia ^[87]	Valid	Gen. et Sp. nov.	Mayr & Manegold	Rupelian		Germany	An early Passeriformes Incertae Sedis, type species is Wi. weissi

Newly named pterosaurs[edit]

Name	Status	Authors	Age	Unit	Location	Notes
Cathayopterus^[88]	Valid	Wang Zhou	Early Cretaceous	Yixian Formation	China	Apterodactyloid. The type species is Cathayopterus grabaui.
Caviramus^[89]	Valid	Fröbisch Fröbisch	Late Triassic	Kössen Formation	Switzerland	Arhamphorhynchoid. The type species is Caviramus schesaplanensis.
Longchengpterus^[90]	Valid	Wang Li Duan Cheng	Early Cretaceous	Jiufotang Formation	China	A pterodactyloid. The type species is Longchengpterus zhoai.
Muzquizopteryx^[91]	Valid	Frey Buchy Stinnesbeck Gonzalez Stefano	Late Cretaceous	Austin Group	Mexico	The type species is Muzquizopteryx coahuilensis.
Yixianopterus^[92]	Valid	Lü Ji Yuan Gao Sun Ji	Early Cretaceous	Yixian Formation	China	A pterodactyloid. The type species is Yixianopterus jingangshanensis.

Synapsids[edit]

Non-mammalian[edit]

Name	Status	Authors	Age	Unit	Location	Notes
Herpetoskylax^[93]	Valid	Sidor Rubidge	Late Permian		South Africa	The type species is Herpetoskylax hopsoni.
Elliotherium^[94]	Valid	Sidor Hancox	Upper Triassic	Lower Elliot Formation	South Africa	The type species is Elliotherium kerstenae (emmend.).
Idelesaurus^[95]	Valid	Kurkin	Upper Permian		Russia	The type species is Idelisaurus tataricus.
Langbergia^[96]	Valid	Abdala Neveling Welman	Early Triassic	Burgersdorp Formation	South Africa	The type species is Langbergia modisei.
Nanogomphodon^[97]	Valid	Hopson Sues	Middle Triassic	Erfurt Formation	Germany	The type species is Nanogomphodon wildi.
Oromycter^[98]	Valid	Reisz	Early Permian		USA	The type species is Oromycter dolesorum.
Pachydectes^[99]	Valid	Rubidge Sidor Modesto	Middle Permian	Koonap Formation	South Africa	The type species is Pachydectes elsi.
Paraburnetia^[100]	Valid	Smith Rubidge Sidor	Upper Permian	Teekloof Formation	South Africa	The type species is Paraburnetia sneeubergensis.
Protheriodon^[101]	Valid	Bonaparte, Soares, & Schultz	Middle Triassic	Santa Maria Formation	Brazil	Type sp P. estudianti.

Mammals[edit]

Name	Status	Authors	Age	Unit	Location	Notes
Anthracoxyaena^[102]	Disputed	Tong & Wang	Early Eocene	Wutu Formation	China	A member of the family Oxyaenidae. Genus includes new species A. palustris. Subsequently, Solé, Gheerbrant & Godinot (2013) considered the genus Anthracoxyaena to be a junior synonym of the genus Arfia, though the authors maintained A. palustris as a distinct species within the latter genus.^[103]
Asiohyopsodus^[102]	Valid	Tong & Wang	Early Eocene	Wutu Formation	China	A member of the family Hyopsodontidae. Genus includes new species A. confuciusi.
Asioictops^[102]	Valid	Tong & Wang	Early Eocene	Wutu Formation	China	A member of the family Leptictidae. Genus includes new species A. mckennai.
Celaenolambda^[102]	Valid	Tong & Wang	Early Eocene	Wutu Formation	China	A possible member of the family Archaeolambdidae. Genus includes new species C. wangzhaoi.
Chowliia^[102]	Valid	Tong & Wang	Early Eocene	Wutu Formation	China	A member of the family Isectolophidae. Genus includes new species C. laoshanensis.
Dianomomys^[102]	Valid	Tong & Wang	Early Eocene	Wutu Formation	China	A possible member of the family Paromomyidae. Genus includes new species D. ambiguus.
Dissacus bohaiensis^[102]	Valid	Tong & Wang	Early Eocene	Wutu Formation	China	A member of Mesonychia belonging to the family Mesonychidae.
Hapalodectes huanghaiensis^[102]	Valid	Tong & Wang	Early Eocene	Wutu Formation	China	A member of Mesonychia belonging to the family Hapalodectidae.
Heterocoryphodon？yuntongi^[102]	Valid	Tong & Wang	Early Eocene	Wutu Formation	China	A member of the family Coryphodontidae.
Hylomysoides^[102]	Valid	Tong & Wang	Early Eocene	Wutu Formation	China	A possible member of the family Amphilemuridae. Genus includes new species H. qiensis.
Luchenus^[102]	Valid	Tong & Wang	Early Eocene	Wutu Formation	China	A member of the family Erinaceidae. Genus includes new species L. erinaceanus.
Migrostylops^[102]	Valid	Tong & Wang	Early Eocene	Wutu Formation	China	A member of Arctostylopida belonging to the family Arctostylopidae. Genus includes new species M. roboreus and M. rosella.
Pappomoropus^[102]	Valid	Tong & Wang	Early Eocene	Wutu Formation	China	A member of the family Eomoropidae. Genus includes new species P. taishanensis.
Paresthonyx^[102]	Valid	Tong & Wang	Early Eocene	Wutu Formation	China	A member of Tillodontia. Genus includes new species P. orientalis.
Preonictis^[102]	Valid	Tong & Wang	Early Eocene	Wutu Formation	China	A member of the family Hyaenodontidae belonging to the subfamily Proviverrinae. Genus includes new species P. youngi.
Qilulestes^[102]	Valid	Tong & Wang	Early Eocene	Wutu Formation	China	A possible member of the family Amphilemuridae. Genus includes new species Q. schieboutae.
Scileptictis^[102]	Valid	Tong & Wang	Early Eocene	Wutu Formation	China	A member of the family Leptictidae. Genus includes new species S. simplus and S? stenotalus.
Suyinia^[102]	Valid	Tong & Wang	Early Eocene	Wutu Formation	China	A member of Anagalida belonging to the family Pseudictopidae. Genus includes new species S. changleensis.
Taishanomys parvulus^[102]	Valid	Tong & Wang	Early Eocene	Wutu Formation	China	Arodent.
Talpilestes^[102]	Valid	Tong & Wang	Early Eocene	Wutu Formation	China	A possible member of the family Nyctitheriidae. Genus includes new species T. asiatica.
Variviverra^[102]	Valid	Tong & Wang	Early Eocene	Wutu Formation	China	A member of the family Viverravidae. Genus includes new species V. vegetatus.
Wutucoryphodon^[102]	Valid	Tong & Wang	Early Eocene	Wutu Formation	China	A member of the family Coryphodontidae. Genus includes new species W. xianwui.
Wutuhyus^[102]	Valid	Tong & Wang	Early Eocene	Wutu Formation	China	A member of Suiformes of uncertain phylogenetic placement. Genus includes new species W. primiveris.
Yupingale^[102]	Valid	Tong & Wang	Early Eocene	Wutu Formation	China	A member of the family Astigalidae. Genus includes new species Y. weifangensis.
Zodiocyon^[102]	Valid	Tong & Wang	Early Eocene	Wutu Formation	China	A member of the family Miacidae. Genus includes new species Z. zetesios.

Trace fossils[edit]

The trace fossil genera Nihilichnus (Nihilichnus nihilicus and Nihilichnus mortalis), Machichnus (Machichnus regularis, Machichnus multilineatus, and Machichnus bohemicus) and Brutalichnus (Brutalichnus brutalis) are described from bite traces.^[104]

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^ ^a ^b ^c ^d ^e ^f ^g ^h ⁱ ^j ^k ^l ^m ⁿ ^o ^p ^q ^r ^s ^t ^u ^v ^w ^x ^y Yongsheng Tong; Yingwen Wang (2006). "Fossil mammals from the early Eocene Wutu Formation of Shandong Province". Palaeontologia Sinica. 192, New Series C28: 1–195. ISBN 7-03-016894-1.

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