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Contents

   



(Top)
 


1 Shell description  



1.1  The clausilium  







2 Anatomy  





3 Taxonomy  



3.1  2005 taxonomy  







4 Genera  



4.1  Clausiliinae  





4.2  Alopiinae A. J. Wagner, 1913  





4.3  Constrictinae  





4.4  Eualopiinae  





4.5  Garnieriinae  





4.6  Laminiferinae  





4.7  Mentissoideinae  





4.8  Neniinae Wenz, 1923  





4.9  Peruiniinae H. Nordsieck, 2005  





4.10  Phaedusinae A. J. Wagner, 1922  







5 Synonyms  





6 Conservation  





7 References  





8 Further reading  





9 External links  














Clausiliidae






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From Wikipedia, the free encyclopedia
 


Clausiliidae
Cochlodina laminata
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Mollusca
Class: Gastropoda
Subclass: Heterobranchia
Order: Stylommatophora
Suborder: Helicina
Infraorder: Clausilioidei
Superfamily: Clausilioidea
Family: Clausiliidae
J. E. Gray, 1855[1]
Type genus
Clausilia

Draparnaud, 1805

Clausiliidae, also known by the common name door snails, is a taxonomic family of small, very elongate, mostly left-handed, air-breathing land snails, sinistral terrestrial pulmonate gastropod mollusks.[2]

With over 1700 recognized recent and fossil species,[2] this is among the most diverse families of terrestrial gastropods (cf. Orthalicidae), although the marine gastropod family Pyramidellidae is larger.

Most species of Clausiliidae have an anatomical structure known as a clausilium, which enables the snail to close off the aperture of the shell with a sliding "door".

Shell description[edit]

Megalophaedusa martensi is the largest species of the family Clausiliidae in the world. The individual in the photograph has a shell which is 45.5 mm long. The whole snail weighs 3.2 g live.

Almost all the species of snails in the family of door snails are left-handed, which is an uncommon feature in gastropod shells in general.

These snails have shells which are extremely high-spired, with numerous whorls.

The shells tend to be club-shaped, tapering at both ends to a rounded nub. The aperture usually has visible folds.

The clausilium[edit]

The clausiliumofClausilia dubia

Clausiliids are also very unusual among pulmonate gastropods in that most of them have a "door" or clausilium. The clausilium is not the same thing as an operculum, which does not exist at all in pulmonate gastropods.

The clausilium is a calcareous structure, tongue-shaped or spoon-shaped, which can close the aperture of the snail shell to protect the soft parts against predation by animals such as carnivorous beetle larvae. The narrow end of the clausilium slides in the grooves that are formed by the folds on the inside of the shell.

Anatomy[edit]

In this family, the number of haploid chromosomes lies between 21 and 30 (according to the values in this table).[3]

Taxonomy[edit]

The type genusisClausilia Draparnaud, 1805.

The family Clausiliidae is classified within the informal group Sigmurethra, itself belonging to the clade Stylommatophora within the clade Eupulmonata (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005).[4]

2005 taxonomy[edit]

The taxonomy of the Gastropoda by Bouchet & Rocroi, 2005 recognizes subfamilies as follows:

subfamily Clausiliinae Gray, 1855

subfamily Alopiinae A. J. Wagner, 1913

subfamily † Constrictinae H. Nordsieck, 1981[5]

subfamily Garnieriinae C. Boettger, 1926

subfamily † Eualopiinae H. Nordsieck, 1978

subfamily Laminiferinae Wenz, 1923

subfamily Mentissoideinae Lindholm, 1924

subfamily Neniinae Wenz, 1923 - Neniastrinae H. B. Baker, 1930

subfamily Phaedusinae A. J. Wagner, 1922

subfamily Serrulininae Ehrmann, 1927

Genera[edit]

Genera include:

Clausiliinae[edit]

Tribe Acrotomini H. Nordsieck, 1979

tribe Baleini

Tribe Boettgeriini H. Nordsieck, 1979

tribe Clausiliini

Tribe † Emarginariini H. Nordsieck, 2007

Tribe Euxinellini Neubert, 2002

Tribe Filosini H. Nordsieck, 1979

tribe Gracillariini H. Nordsieck, 1979

Tribe Mentissoideini Lindholm, 1924

Tribe Olympicolini Neubert, 2002

Tribe Strigileuxinini H. Nordsieck, 1994

Tribe Strumosini H. Nordsieck, 1994

Alopiinae A. J. Wagner, 1913[edit]

Alopiinae[11]

tribe Alopiini

tribe Cochlodinini Lindholm, 1925 (1923)

tribe Delimini R. Brandt, 1956

tribe Medorini H. Nordsieck, 1997

Constrictinae[edit]

Fossil subfamily Constrictinae contains genera:

Eualopiinae[edit]

Fossil subfamily Eualopiinae contains genera:

Tribe † Eualopiini H. Nordsieck, 1978

Tribe † Rillyini † H. Nordsieck, 1985

Garnieriinae[edit]

Laminiferinae[edit]

Tribe Laminiferini Wenz, 1923

Tribe † Oospiroidesini H. Nordsieck, 2007

Tribe † Polloneriini H. Nordsieck, 2007

Mentissoideinae[edit]

synonym of the tribe Mentissoideini Lindholm, 1924 (superseded classification)

Neniinae Wenz, 1923[edit]

Tribe Neniini Wenz, 1923

Peruiniinae H. Nordsieck, 2005[edit]

Phaedusinae A. J. Wagner, 1922[edit]

Tribe † Disjunctariini H. Nordsieck, 2014

Tribe † Nordsieckiini H. Nordsieck, 2007

tribe Phaedusini A. J. Wagner, 1922

Tribe † Serrulellini H. Nordsieck, 2007

Tribe Serrulinini Ehrmann, 1927

Tribe Synprosphymini H. Nordsieck, 2007

Synonyms[edit]

Conservation[edit]

Although non-marine molluscs appear to be exceptionally vulnerable to extinction,[16] the IUCN Red list listed only 9 species[17] from this family.

References[edit]

  1. ^ J. E. Gray. April 14, 1855. Catalogue of Pulmonata or air-breathing Mollusca in the collection of the British Museum, Part I: 156.
  • ^ a b MolluscaBase eds. (2023). MolluscaBase. Clausiliidae L. Pfeiffer & J. E. Gray, 1855. Accessed through: World Register of Marine Species at: https://www.molluscabase.org/aphia.php?p=taxdetails&id=833936 on 2023-04-11
  • ^ Barker G. M.: Gastropods on Land: Phylogeny, Diversity and Adaptive Morphology. in Barker G. M. (ed.): The biology of terrestrial molluscs. CABI Publishing, Oxon, UK, 2001, ISBN 0-85199-318-4. 1-146, cited pages: 139 and 142.
  • ^ a b c d e f Bouchet, Philippe; Rocroi, Jean-Pierre; Frýda, Jiri; Hausdorf, Bernard; Ponder, Winston; Valdés, Ángel & Warén, Anders (2005). "Classification and nomenclator of gastropod families". Malacologia. 47 (1–2). Hackenheim, Germany: ConchBooks: 1–397. ISBN 3-925919-72-4. ISSN 0076-2997.
  • ^ Nordsieck H. (1981). Archiv für Molluskenkunde 111(1-3): 101.
  • ^ Nordsieck H. (1978). Archiv für Molluskenkunde 109(1-3): 104.
  • ^ Nordsieck H. (1985). Heldia 1(3): 83.
  • ^ Rodrigo B. Salvador; Abraham S.H. Breure (2020). "Type material of Clausiliidae door snails from Philippe Dautzenberg in the collection of the Museum of New Zealand Te Papa Tongarewa". Tuhinga: Records of the Museum of New Zealand Te Papa Tongarewa. 31. Te Papa: 56–69. ISSN 1173-4337. Wikidata Q106839645.
  • ^ a b c d e f g h i "Clausiliini". Fauna Europaea, last update 27 January 2011, accessed 27 April 2011.
  • ^ "Graciliaria". Fauna Europaea, last update 27 January 2011, accessed 27 April 2011.
  • ^ Uit de Weerd D. R. (2004). "Molecular phylogenetic history of eastern Mediterranean Alopiinae, a group of morphologically indeterminate land snails". Doctoral thesis, Leiden University, ISBN 90-6464-874-3.
  • ^ Gittenberger E. & Uit de Weerd D. R. (2009). "Summarizing data on the Inchoatia taxa, including Inchoatia megdova bruggeni subspec. nov. (Gastropoda, Pulmonata, Clausiliidae)" Zoologische Mededelingen 83 http://www.zoologischemededelingen.nl/83/nr03/a08 Archived 16 January 2014 at the Wayback Machine
  • ^ Rodrigo B. Salvador; Alina Wahab; Nicole E. Phillips; Abraham S.H. Breure (2021). "South American and Trinidadian terrestrial Gastropoda in the collection of the Museum of New Zealand Te Papa Tongarewa". Tuhinga: Records of the Museum of New Zealand Te Papa Tongarewa. 32. Te Papa: 64–80. ISSN 1173-4337. Wikidata Q116264204.
  • ^ cf. Nordsieck H. (2006). "Species list of recent Clausiliidae".
  • ^ a b Maassen W. J. M. & Gittenberger E. (2007). "Three new clausiliid land snails from Tonkin, northern Vietnam (Gastropoda: Pulmonata: Clausiliidae)". Zoologische Mededelingen 81(1): http://www.zoologischemededelingen.nl/81/nr01/a10 Archived 7 October 2011 at the Wayback Machine
  • ^ Lydeard, C.; Cowie R.; Ponder, W.F.; et al. (April 2004). "The global decline of nonmarine mollusks". BioScience. 54 (4): 321–330. doi:10.1641/0006-3568(2004)054[0321:TGDONM]2.0.CO;2.
  • ^ IUCN 2009. IUCN Red List of Threatened Species. Version 2009.2. <www.iucnredlist.org>. Downloaded on 5 March 2010.
  • Further reading[edit]

    External links[edit]


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