Contents

Geomitridae

Geomitridae
Cernuella virgata
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Mollusca
Class:	Gastropoda
Subclass:	Heterobranchia
Order:	Stylommatophora
Superfamily:	Helicoidea
Family:	Geomitridae Boettger, 1909
Type genus
Geomitra Swainson, 1840
Genera
Actinella Backeljaia Candidula Caseolus Cernuella Cernuellopsis Cochlicella Discula Geomitra Helicella Helicopsis Microxeromagna Orexana Plentuisa Ponentina Trochoidea Xerocrassa Xerogyra Xerolenta Xeroplexa Xerosecta Xerotricha Zarateana

Geomitridae is a taxonomic family of small to medium-sized air-breathing land snails, terrestrial pulmonate gastropod mollusks in the superfamily Helicoidea.^[1][2]

The family is characterized by the presence of a free right ommatophoral retractor (passing outside the peni-oviducal angle) and a double stimulatory apparatus.^[1] A free right ommatophoral retractor has been linked to adaptation to xeric habitats^[3]

The family Geomitridae is subdivided in the following subfamilies (according to Razkin et al., 2015^[1])

• Subfamily Geomitrinae
  • tribe Cochlicellini Schileyko, 1972
  • tribe Geomitrini C. Boettger, 1909 - synonym: Ochthephilinae Zilch, 1960 (n.a.)
  • tribe Ponentinini Razkin et al., 2015
• Subfamily Helicellinae
  • tribe Cernuellini Schileyko, 1991
  • tribe Helicellini Ihering, 1909 - synonym: Jacostidae Pilsbry, 1948 (inv.)
  • tribe Helicopsini H. Nordsieck, 1987
  • tribe Plentuisini Razkin et al., 2015
  • tribe Trochoideini H. Nordsieck, 1987

Genera with the family Geomitridae include:^[1][4]

Not belonging to a subfamily:

• Keraea Gude, 1911
• Peridotitea Torres Alba, Holyoak, D. T., Holyoak, G. A., Vázquez Toro & Ripoll, 2018
• Subfamily Geomitrinae
• tribe Cochlicellini
  • Cochlicella Férussac, 1821
  • Monilearia Mousson, 1872
  • Obelus W. Hartmann, 1842
  • Ripkeniella Hutterer & E. Gittenberger, 1998
• tribe Geomitrini
  • Actinella R. T. Lowe, 1852^[5]
  • Callina R. T. Lowe, 1855
  • Caseolus R. T. Lowe, 1852^[5]
  • Discula R. T. Lowe, 1852^[5]
  • Disculella Pilsbry, 1895^[5]
  • Domunculifex Brozzo, De Mattia, Harl & Neiber, 2020
  • Geomitra Swainson, 1840^[5]
  • Helicomela R. T. Lowe, 1855
  • Heterostoma W. Hartmann, 1843:^[5] synonym of Steenbergia Mandahl-Barth, 1950 (invalid: junior homonym of Heterostoma de Filippi, 1837 [Platyhelminthes])
  • Hystricella R. T. Lowe, 1855^[5]
  • Lemniscia R. T. Lowe, 1855^[5]
  • Moreletina de Frias Martins, 2002^[5]
  • Plebecula R. T. Lowe, 1852
  • Pseudocampylaea L. Pfeiffer, 1877^[5]
  • Serratorotula Groh & Hemmen, 1986^[5]
  • Spirorbula R. T. Lowe, 1852^[5]
  • Steenbergia Mandahl-Barth, 1950
  • Testudodiscula Brozzo, De Mattia, Harl & Neiber, 2020
  • Wollastonaria De Mattia, Neiber & Groh, 2018
• tribe Ponentinini
  • Ponentina P. Hesse, 1921

subfamily Helicellinae

• tribe Cernuellini
  • Alteniella Clerx & Gittenberger, 1977
  • Cernuella Schlüter, 1838
  • Microxeromagna Ortiz de Zárate López, 1950
  • Xeroplana Monterosato, 1892
  • Xerosecta Monterosato, 1892
• tribe Helicellini
  • Backeljaia Chueca et al., 2018
  • Candidula Kobelt, 1871
  • Helicella Férussac, 1821
  • Orexana Chueca et al., 2018
  • Xerogyra Monterosato, 1892
  • Xeroleuca Kobelt, 1877
  • Xeroplexa Monterosato, 1892
  • Xerotricha Monterosato, 1892
  • Zarateana Chueca et al., 2018
• tribe Helicopsini H. Nordsieck, 1987
  • Helicopsis Fitzinger, 1833
  • Pseudoxerophila Westerlund, 1879
  • Xerolenta Monterosato, 1892
  • Xeromunda Monterosato, 1892
  • Xeropicta Monterosato, 1892
• tribe Plentuisini
  • Plentuisa Puente & Prieto, 1992
• tribe Trochoideini
  • Trochoidea T. Brown, 1827
  • Xerocrassa Monterosato, 1892

• Bouchet P., Rocroi J.P., Hausdorf B., Kaim A., Kano Y., Nützel A., Parkhaev P., Schrödl M. & Strong E.E. (2017). Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia. 61(1-2): 1-526

Wikimedia Commons has media related to Geomitridae.

• https://liferecovernatura.madeira.gov.pt
• Razkin, O., Gómez-Moliner, B. J., Prieto, C. E., Martínez-Ortí, A., Arrébola, J. R., Muñoz, B., Chueca, L. J. & Madeira, M. J. (2015). Molecular phylogeny of the western Palaearctic Helicoidea (Gastropoda, Stylommatophora). Molecular Phylogenetics and Evolution. 83: 99-117