Jump to content
 







Main menu
   


Navigation  



Main page
Contents
Current events
Random article
About Wikipedia
Contact us
Donate
 




Contribute  



Help
Learn to edit
Community portal
Recent changes
Upload file
 








Search  

































Create account

Log in
 









Create account
 Log in
 




Pages for logged out editors learn more  



Contributions
Talk
 



















Contents

   



(Top)
 


1 Phylogeny  





2 Characteristics  





3 Distribution and habitat  





4 Behaviour and ecology  





5 Threats  



5.1  Habitat degradation  





5.2  Exploitation and fragmentation  







6 Conservation  



6.1  Priority projects  







7 Cultural references  





8 References  





9 External links  














Siamese crocodile






العربية
Azərbaycanca
Беларуская
Български
Brezhoneg
Català
Чӑвашла
Cebuano
Čeština
Deutsch
Diné bizaad
Español
Esperanto
Euskara
فارسی
Français
Galego

Հայերեն
Hrvatski
Bahasa Indonesia
Italiano
עברית
Kaszëbsczi
Lietuvių
Magyar
Македонски
مصرى
Bahasa Melayu
Nederlands

پنجابی
Polski
Português
Русский
Simple English
Slovenčina
Српски / srpski
Suomi
Svenska
ி

Türkçe
Tiếng Vit
Winaray


 

Edit links
 









Article
Talk
 

















Read
Edit
View history
 








Tools
   


Actions  



Read
Edit
View history
 




General  



What links here
Related changes
Upload file
Special pages
Permanent link
Page information
Cite this page
Get shortened URL
Download QR code
Wikidata item
 




Print/export  



Download as PDF
Printable version
 




In other projects  



Wikimedia Commons
Wikispecies
 
















Appearance
   

 






From Wikipedia, the free encyclopedia
 


Siamese crocodile
Temporal range: PleistocenePresent, 2.6–0 Ma[1]

O

S

D

C

P

T

J

K

Pg

N

Siamese crocodile at Moscow Zoo

Conservation status


Critically Endangered  (IUCN 3.1)[2]

CITES Appendix I (CITES)[2]

Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Clade: Archosauromorpha
Clade: Archosauriformes
Order: Crocodilia
Family: Crocodylidae
Genus: Crocodylus
Species:
C. siamensis
Binomial name
Crocodylus siamensis

Schneider, 1801

Synonyms
  • Crocodylus ossifragus Dubois, 1908

The Siamese crocodile (Crocodylus siamensis) is a medium-sized freshwater crocodile native to Indonesia (Borneo and possibly Java), Brunei, East Malaysia, Laos, Cambodia, Myanmar, Thailand and Vietnam. The species is critically endangered and already extirpated from many regions. Its other common names include Siamese freshwater crocodile, Singapore small-grain, and soft-belly.[3]

Phylogeny

[edit]
Fossils, formerly referred to as Crocodylus ossifragus

Below is a cladogram based on a 2018 tip dating study by Lee & Yates simultaneously using morphological, molecular (DNA sequencing), and stratigraphic (fossil age) data,[4] as revised by the 2021 Hekkala et al. paleogenomics study using DNA extracted from the extinct Voay.[5]

Crocodylinae

Voay

Crocodylus
Asia+Australia

Crocodylus johnstoni Freshwater crocodile

Crocodylus novaeguineae New Guinea crocodile

Crocodylus mindorensis Philippine crocodile

Crocodylus porosus Saltwater crocodile

Crocodylus siamensis Siamese crocodile

Crocodylus palustris Mugger crocodile

Africa+New World

Crocodylus suchus West African crocodile

Crocodylus niloticus Nile crocodile

New World

Crocodylus moreletii Morelet's crocodile

Crocodylus rhombifer Cuban crocodile

Crocodylus intermedius Orinoco crocodile

Crocodylus acutus American crocodile

Characteristics

[edit]
Siamese Crocodile – Biblical Zoo

The Siamese crocodile is a medium-sized, freshwater crocodilian, with a relatively broad, smooth snout and an elevated, bony crest behind each eye. Overall, it is olive-green, with some variation to dark-green.[6] Young individuals measure 1.2 to 1.5 m (3 ft 11 in – 4 ft 11 in) and weigh 6 to 12 kg (13 to 26 lb), growing to a length of 2.1 to 3 m (6 ft 11 in – 9 ft 10 in) and a weight of 40 to 120 kg (88 to 265 lb) as an adult.[7][8] Three individuals measuring 2.12 to 2.63 m (6 ft 11 in – 8 ft 8 in) and weighing 40–87 kg (88–192 lb) had bite force of 2,073–4,577 N (466–1,029 lbf).[9] Large male individuals reach 4 m (13 ft 1 in) and 350 kg (770 lb) in weight.[10]

Distribution and habitat

[edit]

Siamese crocodiles occur in a wide range of freshwater habitats, including slow-moving rivers and streams, lakes, seasonal oxbow lakes, marshes and swamps.[11]

Behaviour and ecology

[edit]
Siamese crocodile sleeping with its mouth open to release heat

Despite conservation concerns, many aspects of C. siamensis life history in the wild remain unknown, particularly regarding its reproductive biology.[11]

Adults feed mainly on fish and snakes, but also eat amphibians and small mammals.[3]

Very little is known about the natural history of this species in the wild, but females build mound-nests constructed from scraped-up plant debris mixed with mud.[6] In captivity, these crocodiles breed during the wet season (April to May), laying between 15 and 50 eggs, which are then guarded until they hatch.[12] After incubation, the female will assist her young as they break out of their eggs and then carry the hatchlings to the water in her jaws.[13]

Pure, unhybridised examples of this species are generally unaggressive towards humans,[14] and there are only four confirmed attacks, none of them fatal. One was defending its young,[15] another was probably defending itself,[16] one was provoked,[17] and the reason for the last is unclear.[18] A fifth attack in 1928 that was probably done by a Siamese crocodile was fatal, with the victim being a child.[19]

Threats

[edit]

Siamese crocodiles are under threat from human disturbance and habitat occupation, which is forcing remaining populations to the edges of their former range.[12] Extinct from 99% of its original range, the Siamese crocodile is considered one of the least studied and most critically endangered crocodilians in the world.[11] Although few wild populations remain, more than 700,000 C. siamensis are held on commercial crocodile farms in Southeast Asia.[11]

In 1992, it was believed to be extremely close to or fully extinct in the wild until 2000 when scientists from Fauna and Flora International and the Government of Cambodia's Forestry Administration confirmed the presence of Siamese crocodiles in the Cardamom Mountains in Southwest Cambodia.[20][21] Since then, surveys have identified around 30 sites in Cambodia that contain wild Siamese crocodiles (conservatively estimated to number between 200 and 400 individuals in total), a tiny population in Thailand (possibly as few as two individuals, discounting recent reintroductions), a small population in Vietnam (possibly fewer than 100 individuals), Cat Tien National Park has about 200, and a more sizeable population in Laos.[22] In March 2005, conservationists found a nest containing juvenile Siamese crocodiles in the southern Lao province of Savannakhet.[23] There are no recent records from Malaysia, BurmaorBrunei. A small but important population of the crocodiles is also known to live in East Kalimantan, Indonesia.[24][25]

Habitat degradation

[edit]

Factors causing loss of habitat include: conversion of wetlands for agriculture, chemical fertilisers use, use of pesticides in rice production, and an increase in the population of cattle.[26] The effects of warfare stemming from the conflicts in Vietnam, Laos, and Cambodia during the Vietnam War (from land mines to aerial bombardment) have also been factored.[27]

Many river systems, including those in protected areas, have hydroelectric power dams approved or proposed, which are likely to cause the loss of about half of the remaining breeding colonies within the next ten years.[11] One cause for habitat degradation via hydrological changes, for the Siamese crocodile, is the implementation of dams on the upper Mekong River and its major tributaries.[28] Potential impacts of dam construction include wetland loss and altered flooding cycle with a dry season flow 50% greater than under natural conditions.[29]

Exploitation and fragmentation

[edit]
Siamese crocodile farm on Tonle Sap in Cambodia

Illegal capture of wild crocodiles for supply to farms is an ongoing threat, as well as incidental capture/drowning in fishing nets and traps.[11] The Siamese crocodile currently has extremely low and fragmented remaining populations with little proven reproduction in the wild.[30]

Siamese crocodiles have historically been captured for skins and to stock commercial crocodile farms. In 1945, skin hunting for commercial farms was banned by the French colonial administration of Cambodia.[31] In the late 1940s, populations spurred the development of farms and harvesting wild crocodiles for stocking these farms.[32] Protection was abolished by the Khmer Rouge (1975–79) but later reinstated under Article 18 of the Fishery Law of 1987, which "forbids the catching, selling, and transportation of...[wild] crocodiles..."[31]

Crocodile farming now has a huge economic impact in the provinces surrounding Tonle Sap, where 396 farms held over 20,000 crocodiles in 1998.[31] Also, many crocodiles were exported from Cambodia since the mid-1980s to stock commercial farms in Thailand, Vietnam, and China.[33]

Despite legal protection, a profitable market exists for the capture and sale of crocodiles to farms since the early 1980s.[33] Chronic hunting has led to the decline of the wild Siamese crocodile.[22][30]

Conservation

[edit]

This crocodile is classified as critically endangered on the IUCN Red List, and is listed on Appendix I of CITES.[2] It is one of the most endangered crocodiles in the wild, although it is extensively bred in captivity.[12]

Yayasan Ulin (The Ironwood Foundation) ran a small project to conserve an important wetland habitat in the area of East Kalimantan which is known to contain the crocodiles.[34]

In Cambodia Fauna and Flora International and the Government of Cambodia's Forestry Administration have established the Cambodian Crocodile Conservation Programme for the protection and recovery of Siamese crocodiles. This programme works with a network of indigenous villages who are helping to protect key sites such as Veal Veng Marsh (Veal Veng District), the Tatai River (Thmar Bang District) and the Araeng River. The latter is considered to have the second largest population of Siamese crocodiles in the world, but is currently threatened by the proposed construction of a massive dam in the river. During the heavy monsoon period of June–November, Siamese crocodiles take advantage of the increase in water levels to move out of the river and onto large lakes and other local bodies of water, returning to their original habitat once water levels start receding back to their usual levels.[35]

The Cambodian Crocodile Conservation Programme conducted DNA analysis of 69 crocodiles in Phnom Tamao Wildlife Rescue Centre in Cambodia 2009, and found 35 of them were purebred C. siamensis. Conservationists from the Forestry Administration and Fauna and Flora International subsequently launched a conservation breeding program at the Centre. Since 2012, approximately 50 purebred Cambodian Siamese crocodiles have been released into community-protected areas to reinforce the depleted wild populations.[36]

Poaching is a severe threat to this species, with the value of wild Siamese crocodiles reaching hundreds of dollars in the black market, where they are illegally taken into crocodile farms and hybridized with other, larger species.[37] The total wild population is unknown, since most groups are in isolated areas where access is extremely complicated. A number of captively held individuals are the result of hybridization with the saltwater crocodile, but several thousand "pure" individuals do exist in captivity, and are regularly bred at crocodile farms, especially in Thailand.

Pang Sida National Park in Thailand, near Cambodia, has a project to reintroduce Siamese crocodile into the wild. A number of young crocodiles have been released into a small and remote river in the park, not accessible to visitors.[22]

The Wildlife Conservation Society (WCS) is working with the government of Laos on a new programme to save this critically endangered crocodile and its wetland habitat. In August, 2011, a press release announced the successful hatching of a clutch of 20 Siamese crocodiles. These eggs were then incubated at the Laos Zoo. This project represents a new effort by WCS to conserve the biodiversity and habitat of Laos’ Savannakhet Province, promotes conservation of biodiversity for the whole landscape, and relies on community involvement from local residents.[38]

In September 2021, eight hatchlings were found in a wildlife sanctuary in eastern Cambodia.[39]

InCambodia's Southern Cardamom National Park, sixty Siamese crocodiles from five distinct nests successfully hatched in July 2024. This marks the largest wild breeding record of the species this century and significantly enhances the survival prospects of this critically endangered reptile.[40][41]

Priority projects

[edit]

High priority projects include:[42]

Other projects include:[35]

Cultural references

[edit]

AMalay folktale features a crocodile that is outwitted by a mouse-deer and buffaloes.[43] A Siamese crocodile has been cited in the Thai folkloreofCentral Thailand's Krai Thong ("ไกรทอง") tales have known as well,[44] and was taken to create a television series and movies several times.[45]

In Vietnam a major literary juncture occurred in 1282 when a troublesome crocodile came to inhabit a stretch of the Red River. The scholar Nguyễn Thuyên wrote "Ode to the crocodile" and threw it into the river as an appeal to the beast whereupon the crocodile departed. It was remarkable as historians recorded it as one of the first pieces ever written in the demotic script Chữ Nôm instead of classical Chinese; the emperor amply rewarded the scholar for his literary deed. It marked a shift in Vietnamese literature as the Nôm script gained significant traction as a worthy vehicle for literary works.[46]

A Siamese crocodile stars as the titular monster in the 1978 Thailand film Crocodile.[47]

References

[edit]
  1. ^ Rio, J. P. & Mannion, P. D. (2021). "Phylogenetic analysis of a new morphological dataset elucidates the evolutionary history of Crocodylia and resolves the long-standing gharial problem". PeerJ. 9: e12094. doi:10.7717/peerj.12094. PMC 8428266. PMID 34567843.
  • ^ a b c Bezuijen, M.; Simpson, B.; Behler, N.; Daltry, J. & Tempsiripong, Y. (2012). "Crocodylus siamensis". IUCN Red List of Threatened Species. 2012: e.T5671A3048087. doi:10.2305/IUCN.UK.2012.RLTS.T5671A3048087.en. Retrieved 22 January 2022.
  • ^ a b Crocodilian Species List: (January 2009). "Crocodylus siamensis".{{cite web}}: CS1 maint: numeric names: authors list (link)
  • ^ Michael S. Y. Lee; Adam M. Yates (27 June 2018). "Tip-dating and homoplasy: reconciling the shallow molecular divergences of modern gharials with their long fossil". Proceedings of the Royal Society B. 285 (1881). doi:10.1098/rspb.2018.1071. PMC 6030529. PMID 30051855.
  • ^ Hekkala, E.; Gatesy, J.; Narechania, A.; Meredith, R.; Russello, M.; Aardema, M. L.; Jensen, E.; Montanari, S.; Brochu, C.; Norell, M.; Amato, G. (2021-04-27). "Paleogenomics illuminates the evolutionary history of the extinct Holocene "horned" crocodile of Madagascar, Voay robustus". Communications Biology. 4 (1): 505. doi:10.1038/s42003-021-02017-0. ISSN 2399-3642. PMC 8079395. PMID 33907305.
  • ^ a b Steel, R. (1989). Crocodiles. London.{{cite book}}: CS1 maint: location missing publisher (link)
  • ^ Pye, G. W.; Brown, D. R.; Nogueira, M. F.; Vliet, K. A.; Schoeb, T. R.; Jacobson, E. R. & Bennett, R. A. (2001). "Experimental inoculation of broad-nosed caimans (Caiman latirostris) and Siamese crocodiles (Crocodylus siamensis) with Mycoplasma alligatoris". Journal of Zoo and Wildlife Medicine. 32 (2): 196–201. doi:10.1638/1042-7260(2001)032[0196:eiobnc]2.0.co;2. JSTOR 20096098. PMID 12790420. S2CID 26640928.
  • ^ Whitaker, N. & Sivaraman, C. (2022). "Behavior in a captive family group of Siamese Crocodiles (Crocodylus siamensis) at the Madras Crocodile Bank Trust near Chennai, India". Reptiles & Amphibians. 29 (1): 71–75. doi:10.17161/randa.v29i1.16247. S2CID 245870729.
  • ^ Erickson, G. M.; Gignac, P. M.; Steppan, S. J.; Lappin, A. K.; Vliet, K. A.; Brueggen, J. A.; Inouye, B. D.; Kledzik, D. & Webb, G. J. W. (2012). "Insights into the ecology and evolutionary success of crocodilians revealed through bite-force and tooth-pressure experimentation". PLOS ONE. 7 (3): e31781. Bibcode:2012PLoSO...731781E. doi:10.1371/journal.pone.0031781. PMC 3303775. PMID 22431965.
  • ^ "Farmer raises 6000 crocs". Chinadaily.com.cn. 2012-07-10. Archived from the original on 2016-03-04.
  • ^ a b c d e f Simpson, Boyd; Bezuijen (2010). "Siamese Crocodile Crocodylus Siamensis" (PDF). Crocodiles. Third Edition. Retrieved 2011-11-29.[permanent dead link]
  • ^ a b c Ross, R.P. "Crocodiles: Status Survey and Conservation Action Plan" (Second ed.). Archived from the original on 2011-12-24. Retrieved 2011-11-29.
  • ^ Alderton, D. (1991). Crocodiles and Alligators of the World. Blandford, London.{{cite book}}: CS1 maint: location missing publisher (link)
  • ^ Cox, M.J. van Dijk, P.P, Nabhitabhata, J and Thirakhupt, K. (2009) A photographic guide to Snakes and other reptiles of Thailand and South-East Asia. Asia Books Co. Ltd. Bangkok. ISBN 0883590433.
  • ^ CrocBITE, Worldwide Crocodilian Attack Database: Siamese crocodile, 15 June 2008. Charles Darwin University, Northern Territory, Australia.
  • ^ CrocBITE, Worldwide Crocodilian Attack Database: Siamese crocodile, 15 July 2012. Charles Darwin University, Northern Territory, Australia.
  • ^ CrocBITE, Worldwide Crocodilian Attack Database: Siamese crocodile, 1 January 2017. Charles Darwin University, Northern Territory, Australia.
  • ^ CrocBITE, Worldwide Crocodilian Attack Database: Siamese crocodile, 15 June 1944. Charles Darwin University, Northern Territory, Australia.
  • ^ CrocBITE, Worldwide Crocodilian Attack Database: Siamese crocodile, 1 January 1928. Charles Darwin University, Northern Territory, Australia.
  • ^ University, La Trobe. "Reintroduction of Siamese crocodiles". www.latrobe.edu.au. Retrieved 2024-05-18.
  • ^ Daltry, Jennifer; Han, Sam; Leng, Hor; Ratanapich, Nhek; Piseth, Sorn; Sovannara, Heng; Simpson, Boyd; Starr, Adam; Brook, Sarah; Frechette, Jackson (2015). "Status, distribution and ecology of the Siamese crocodile Crocodylus siamensis in Cambodia". Cambodian Journal of Natural History: 153–164.
  • ^ a b c Lovgren, Stefan (2024-05-18). "They're one of Earth's rarest reptiles. But these crocodiles are bouncing back". National Geographic. Retrieved 2024-05-18.
  • ^ "Endangered crocodiles discovered in Laos". NBC News. 2005-05-20. Retrieved 2024-05-18.
  • ^ "Critically Endangered Siamese Crocodile to Get Key Recognition as Mascot". newsroom.wcs.org. Retrieved 2024-05-18.
  • ^ Kurniati, Hellen- (2007-12-31). "SURVEYS OF SIAMESE CROCODILE (Crocodylus siamensis) HABITAT IN THE MAHAKAM RIVER, EAST KALIMANTAN". Zoo Indonesia (in Indonesian). 16 (2). doi:10.52508/zi.v16i2.116 (inactive 2024-05-18). ISSN 2527-8703.{{cite journal}}: CS1 maint: DOI inactive as of May 2024 (link)
  • ^ Phiapalath, P.; Voladet, H. "Wetland Priority Sites in Lao PDR – the top five priority sites" (PDF). IUCN (Climate Change Impact and Vulnerability Assessment for the Wetlands of the Lower Mekong Basin for Adaptation Planning). Retrieved 2011-11-29.
  • ^ Pierret, Alain; Lacombe, Guillaume; Hoanh, Chu Thai. "Environmental and migratory consequences of the Vietnam War | Environmental Migration Portal". environmentalmigration.iom.int. Retrieved 2024-05-18.
  • ^ Hogan, Z.S.; Moyle, May; Vander Zanden, Baird (2004). "The imperiled giants of the Mekong". American Scientist. 92 (3): 228–237. doi:10.1511/2004.3.228.
  • ^ Lamberts, D (2001). "Tonle Sap fisheries: A Case Study on Floodplain Gillette Fisheries in Siam Reap Cambodia". RAP Publication. Bangkok: FAO Regional Office for Asia and the Pacific.
  • ^ a b Ariyaraphong, Nattakan; Wongloet, Wongsathit; Wattanadilokchatkun, Pish; Panthum, Thitipong; Singchat, Worapong; Thong, Thanyapat; Lisachov, Artem; Ahmad, Syed Farhan; Muangmai, Narongrit; Han, Kyudong; Duengkae, Prateep; Temsiripong, Yosapong; Srikulnath, Kornsorn (2023-03-31). "Should the Identification Guidelines for Siamese Crocodiles Be Revised? Differing Post-Occipital Scute Scale Numbers Show Phenotypic Variation Does Not Result from Hybridization with Saltwater Crocodiles". Biology. 12 (4): 535. doi:10.3390/biology12040535. ISSN 2079-7737. PMC 10136156. PMID 37106736.
  • ^ a b c Thuok; T. (1994). "Country report on crocodile conservation in Cambodia" (PDF). Crocodiles: Proceedings of the 12th Meeting of the Crocodile Specialist Group. IUCN Publications: 3–15.
  • ^ Kimura, W (1969). "Crocodiles in Cambodia, Research Report No. 3 Atawgawa Tropical Garden and Alligator Farm". {{cite journal}}: Cite journal requires |journal= (help)
  • ^ a b Thorbjarnarson, J (2001). "herpetology trip report: Cambodia. Report to Wildlife Conservation Society". {{cite journal}}: Cite journal requires |journal= (help)
  • ^ PAC2013 (2009-07-05). "Siamese Crocodile Conservation Case Study | The Mohamed bin Zayed Species Conservation Fund". Speciesconservation.org. Retrieved 2016-12-15.{{cite web}}: CS1 maint: numeric names: authors list (link)
  • ^ a b "Conserving Siamese crocodiles in Cambodia". Fauna & Flora.
  • ^ "Crocs thought to be extinct found in Cambodia". www.nbcnews.com. 18 November 2009. Retrieved 2022-08-04.
  • ^ Endangered crocodiles hatched in Cambodia. The Associated Press via cbc.ca (June 10, 2010)
  • ^ "WCS Helps Hatch Rare Siamese Crocodiles in Lao PDR". Retrieved 2011-12-01.
  • ^ "Hatchlings of endangered crocodile species found in Cambodia". AP NEWS. 2021-09-21. Retrieved 2021-10-05.
  • ^ "New hope for critically endangered Siamese crocodile". Phys.org.
  • ^ Gaffney, Austyn (July 17, 2024). "For These Nearly Extinct Crocodiles, Life Found a Way". New York Times.
  • ^ Ross, James (1998). "Crocodiles: Status Survey and conservation Action Plan. Second Edition. IUCN/SSC Crocodile Specialist Group". Archived from the original on 2012-03-29. Retrieved 2011-11-29.
  • ^ Winsedtt, R. V. (1906). "Some Mouse-deer Tales". Journal of the Straits Branch of the Royal Asiatic Society. 45: 61–69.
  • ^ "ตำนานและนิทานพื้นบ้านจังหวัดพิจิตร เรื่องชาละวัน- ไกรทอง" (in Thai). ท่องเที่ยวพิจิตร. Archived from the original on 28 September 2016. Retrieved 19 June 2016.
  • ^ "เปิดตัว พญาชาละวัน จากเรื่องไกรทอง 2015 เวอร์ชั่นนี้แซ่บเว่อร์ !!". Kapook.com (in Thai). 28 September 2015. Retrieved 19 June 2016.
  • ^ "Văn tế cá sấu"
  • ^ "Line กนก 8 พฤศจิกายน 2558 FULL". Nation TV (in Thai). 8 November 2015. Archived from the original on 2020-02-16. Retrieved 19 June 2016.
  • [edit]
    Retrieved from "https://en.wikipedia.org/w/index.php?title=Siamese_crocodile&oldid=1235353559"

    Categories: 
    IUCN Red List critically endangered species
    Crocodylidae
    Crocodilians of Asia
    Reptiles of Southeast Asia
    Reptiles of Borneo
    Reptiles of Myanmar
    Reptiles of Cambodia
    Reptiles of India
    Reptiles of Indonesia
    Reptiles of Laos
    Reptiles of Malaysia
    Reptiles of Thailand
    Reptiles of Vietnam
    Critically endangered fauna of Asia
    Reptiles described in 1801
    Apex predators
    Hidden categories: 
    CS1 maint: numeric names: authors list
    CS1 maint: location missing publisher
    CS1: long volume value
    All articles with dead external links
    Articles with dead external links from September 2017
    Articles with permanently dead external links
    CS1 maint: DOI inactive as of May 2024
    CS1 Indonesian-language sources (id)
    CS1 errors: missing periodical
    CS1 Thai-language sources (th)
    Articles with short description
    Short description matches Wikidata
    Articles with 'species' microformats
    Commons category link is on Wikidata
    Webarchive template wayback links
     



    This page was last edited on 18 July 2024, at 21:22 (UTC).

    Text is available under the Creative Commons Attribution-ShareAlike License 4.0; additional terms may apply. By using this site, you agree to the Terms of Use and Privacy Policy. Wikipedia® is a registered trademark of the Wikimedia Foundation, Inc., a non-profit organization.



    Privacy policy

    About Wikipedia

    Disclaimers

    Contact Wikipedia

    Code of Conduct

    Developers

    Statistics

    Cookie statement

    Mobile view



    Wikimedia Foundation
    Powered by MediaWiki