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The clade Australosphenida was proposed by Luo ''et al.'' (2001, 2002) and was initially left unranked, as the authors do not apply the [[Linnaean taxonomy|Linnaean hierarchy]]. In Benton (2005), it is ranked as a '[[Linnaean taxonomy#Taxonomic ranks|superdivision]]', i.e. one or two levels below the [[infraclass]].

The clade Australosphenida was proposed by Luo ''et al.'' (2001, 2002) and was initially left unranked, as the authors do not apply the [[Linnaean taxonomy|Linnaean hierarchy]]. In Benton (2005), it is ranked as a '[[Linnaean taxonomy#Taxonomic ranks|superdivision]]', i.e. one or two levels below the [[infraclass]].



== Evolution ==

== Australosphenid teeth ==

The grouping embodies a hypothesis about the evolution of [[molar (tooth)|molar]] teeth in mammals. Living monotremes are toothless as adults, but the juvenile platypus, fossil monotremes and Ausktribosphenida all share a pattern of three molar [[cusp (dentistry)|cusps]] arranged in a triangle or V shape, which is known as the [[Tribosphenic molar|tribosphenic]] type of molar. Tribosphenic molars have long been held to characterize the subclass [[Theria]] ([[marsupial]]s, [[placental]]s and their extinct relatives), while monotremes were thought to be related to fossil groups with a linear alignment of cusps: [[morganucodon]]tids, [[docodont]]s, [[triconodont]]s and [[multituberculate]]s, all of which were united with the monotremes into the 'subclass [[Prototheria]]'. Defined in this way, the 'Prototheria' is no longer recognised as a valid clade, since the linear cusp pattern is a primitive condition within Mammalia and cannot supply the shared derived character, which is required to establish a subgroup. Instead, the available evidence suggests that the monotremes descend from a [[Mesozoic]] radiation of tribosphenic mammals in the southern continents (hence the name Australosphenida, meaning 'southern wedges'), but this interpretation is highly controversial.

The grouping embodies a hypothesis about the evolution of [[molar (tooth)|molar]] teeth in mammals. Living monotremes are toothless as adults, but the juvenile platypus, fossil monotremes and Ausktribosphenida all share a pattern of three molar [[cusp (dentistry)|cusps]] arranged in a triangle or V shape, which is known as the [[Tribosphenic molar|tribosphenic]] type of molar. Tribosphenic molars have long been held to characterize the subclass [[Theria]] ([[marsupial]]s, [[placental]]s and their extinct relatives), while monotremes were thought to be related to fossil groups with a linear alignment of cusps: [[morganucodon]]tids, [[docodont]]s, [[triconodont]]s and [[multituberculate]]s, all of which were united with the monotremes into the 'subclass [[Prototheria]]'. Defined in this way, the 'Prototheria' is no longer recognised as a valid clade, since the linear cusp pattern is a primitive condition within Mammalia and cannot supply the shared derived character, which is required to establish a subgroup. Instead, the available evidence suggests that the monotremes descend from a [[Mesozoic]] radiation of tribosphenic mammals in the southern continents (hence the name Australosphenida, meaning 'southern wedges'), but this interpretation is highly controversial.



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