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Latest revision Your text
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{{Taxobox

{{Short description|Order of fishes}}

| name = Gonorynchiformes

{{Automatic taxobox

| fossil_range = {{fossil range|143.1|0|[[Berriasian]]–present}}

| fossil_range = Early [[Cretaceous]] - Recent

| image = milkfish.jpg

| image = milkfish.jpg

| image_caption = [[Milkfish]], ''[[Chanos chanos]]''

| image_caption = [[Milkfish]], ''[[Chanos chanos]]''

| regnum = [[Animal]]ia

| taxon = Gonorynchiformes

| authority = [[Lev Berg|L. S. Berg]], 1940

| phylum = [[Chordate|Chordata]]

| classis = [[Actinopterygii]]

| superordo = [[Ostariophysi]]

| ordo = '''Gonorynchiformes'''

| ordo_authority = [[Lev Berg|L. S. Berg]], 1940

| subdivision_ranks = Families

| subdivision_ranks = Families

| subdivision = [[Chanidae]] (milkfish)<br />

| subdivision =

[[Chanidae]] (milkfish)<br>

[[Gonorynchidae]] (beaked salmons)<br />

[[Gonorynchidae]] (beaked salmons)<br>

[[Kneriidae]] (shellears)<br />

[[Kneriidae]] (shellears)<br>

[[Phractolaemidae]] (hingemouths)

[[Phractolaemidae]] (hingemouths)

| type_species = [[Gonorynchus gonorynchus]]

| type_species_authority = ([[Carl Linnaeus|Linnaeus]], 1766)

}}

}}



The '''Gonorynchiformes''' {{IPAc-en|g|ɒ|n|ə|ˈ|r|I|N|k|ᵻ|f|ɔːr|m|iː|z}} are an [[order (biology)|order]] of [[Actinopterygii|ray-finned fish]] that includes the important food source, the [[milkfish]] (''Chanos chanos'', family [[Chanidae]]), and a number of lesser-known types, both [[ocean|marine]] and [[freshwater]].

'''Gonorynchiformes''' is an [[order (biology)|order]] of [[Actinopterygii|ray-finned fish]] that includes the important food source, the [[milkfish]] (''Chanos chanos'', family [[Chanidae]]), and a number of lesser-known types, both [[ocean|marine]] and [[freshwater]].



The alternate spelling "Gonorhynchiformes", with an "h", is frequently seen but not official.

The alternate spelling "Gonorhynchiformes", with an "h", is frequently seen but not official.



Gonorynchiformes have small mouths and no teeth. They are the sole group in the clade Anotophysi, a subgroup of the superorder [[Ostariophysi]]. They are characterized by a primitive [[Weberian apparatus]] formed by the first three vertebrae and one or more cephalic ribs within the head. This apparatus is believed to be a hearing organ, and is found in a more advanced and complex form in the related [[Cypriniformes|cypriniform]] fish, such as [[carp]].<ref name="Nelson">{{cite book | title = Fishes of the World | last = Nelson | first = Joseph, S. | publisher = John Wiley & Sons, Inc. | year = 2006 | isbn = 0-471-25031-7}}</ref> Also like the cypriniforms, the gonorynchiforms produce a substance from their skin when injured that dissolves into the water and acts an alarm signal to other fish.<ref name=EoF>{{cite book |editor=Paxton, J.R. |editor2=Eschmeyer, W.N.|author= Banister, Keith F.|year=1998|title=Encyclopedia of Fishes|publisher= Academic Press|location=San Diego|pages= 96–97|isbn= 0-12-547665-5}}</ref>

Gonorynchiformes have small mouths and no teeth. They are the sole group in the clade Anotophysi, a subgroup of the superorder [[Ostariophysi]]. They are characterized by a primitive [[Weberian apparatus]] formed by the first three vertebrae and one or more cephalic ribs within the head. This apparatus is believed to be a hearing organ, and is found in a more advanced and complex form in the related [[Cypriniformes|cypriniform]] fish, such as [[carp]].<ref name="Nelson">{{cite book | title = Fishes of the World | last = Nelson | first = Joseph, S. | publisher = John Wiley & Sons, Inc. | year = 2006 | isbn = 0-471-25031-7}}</ref> Also like the cypriniforms, the gonorynchiforms produce a substance from their skin when injured that dissolves into the water and acts an alarm signal to other fish.<ref name=EoF>{{cite book |editor=Paxton, J.R. & Eschmeyer, W.N.|author= Banister, Keith F.|year=1998|title=Encyclopedia of Fishes|publisher= Academic Press|location=San Diego|pages= 96–97|isbn= 0-12-547665-5}}</ref>



== Taxonomy ==

== Taxonomy ==

Although many of the families are rather small, there are several fossil genera. This listing of the groups of Gonorynchiformes includes fossil fish with a short description.<ref name="Nelson"/><ref name=":0">{{Citation |last=Taverne |first=Emmanuel Fara, Mireille Gayet, Louis |title=The Fossil Record of Gonorynchiformes |date=2010 |work=Gonorynchiformes and Ostariophysan Relationships |url=https://hal.science/hal-00441945/document |access-date=2024-05-09 |publisher=CRC Press |doi=10.1201/b10194-6/fossil-record-gonorynchiformes-emmanuel-fara-mireille-gayet-louis-taverne |isbn=978-0-429-06156-1}}</ref> They are listed in approximate order of how primitive their characteristics are.

Although many of the families are rather small, there are several fossil genera. Here is a listing of the groups of Gonorynchiformes including fossil fish with a short description.<ref name="Nelson"/> They are listed in approximate order of how primitive their characteristics are.

{{cladogram

|title=Phylogeny of living Gonorynchiformes<ref name="Near, Dornburga & Friedman">Thomas J. Near, Alex Dornburga, Matt Friedman (2014): ''Phylogenetic relationships and timing of diversification in gonorynchiform fishes inferred using nuclear gene DNA sequences (Teleostei: Ostariophysi).'' Molecular Phylogenetics and Evolution, [[DOI:10.1016/j.ympev.2014.07.013]]</ref>

|{{clade|style=font-size:80%;line-height:100%;width:300px;

|label1=Gonorynchiformes

|1={{clade

|label1=Gonorynchidae

|1=''[[Gonorynchus]]''

|2={{clade

|label1=[[Chanidae]]

|1=''[[Chanos (fish)|Chanos]]''

|label2=[[Kneriidae]]

|2={{clade

|label1=Phractolaeminae

|1=''[[Phractolaemus]]''

|label2=Kneriinae

|2={{clade

|1=''[[Kneria]]''

|2={{clade

|1=''[[Parakneria]]''

|label2=Cromeriini

|2={{clade

|1=''[[Cromeria]]''

|2=''[[Grasseichthys]]''

}}

}}

}}

}}

}}

}}

}}

}}



* '''Order''' '''Sorbininardiformes''' Taverne 1999

Order '''Sorbininardiformes''' (extinct)

** '''Family''' '''Sorbininardidae''' Taverne 1999

:: Family '''Sorbininardidae''' (extinct)

*** Genus †''[[Sorbininardus]]'' Taverne 1999

:::: ''[[Sorbininardus]]'' (extinct) - sister group to all other Gonorynchiformes

Order '''Gonorynchiformes'''

* '''Order Gonorynchiformes''' Regan 1909<ref>{{cite web |last1 = Mikko Haaramo |title=Gonorynchiformes |website=Mikko's Phylogeny Archive |access-date= 2013-10-26 |url=http://www.helsinki.fi/~mhaaramo/metazoa/deuterostoma/chordata/actinopterygii/gonorynchiformes/gonorynchiformes.html}}</ref>

: Suborder '''Chanoidei'''

** '''Suborder Gonorynchoidei''' - beaked sandfishes

:::: ''[[Aethalinopsis]]'' (extinct) - sister group to [[Chanidae]] from Early [[Cretaceous]].

*** '''Family''' †'''[[Apulichthyidae]]''' Taverne 1997

**** Genus ''[[Apulichthys]]'' Taverne 1997

:: Family '''[[Chanidae]]''' - milkfishes

*** '''Family [[Gonorynchidae]]''' Fowler 1941

::: Subfamily '''[[Rubiesichthyinae]]''' (extinct)

:::: ''[[Gordichthys]]'' (extinct) - Early Cretaceous Chanid

**** †''[[Charitopsis (fish)|Charitopsis]]'' Gayet 1993 non Trjapitzin 1969

:::: ''[[Rubiesichthys]]'' (extinct) - Early Cretaceous Chanid

**** †''[[Gonorynchidarum]]'' Rana, 1988 otolith

::: Subfamily '''Chaninae'''

**** †''[[Hakeliosomus]]'' Gayet 1993

**** ''[[Ramallichthys]]'' Gayet 1982

:::: ''[[Chanos]]''

:::: ''[[Dastilbe]]'' (extinct) - Early Cretaceous Chanid

**** †''[[Sapperichthys]]'' Amaral, Alvardo-Ortega & Brito, 2013

:::: ''[[Parachanos]]'' (extinct) - Early Cretaceous Chanid

**** Subfamily †Charitosominae

:::: ''[[Tharrhias]]'' (extinct) - Early Cretaceous Chanid

***** †''[[Charitosomus]]'' Hosius & Von Der Marck 1885 - Cretaceous

: Suborder '''Gonorynchoidei''' - beaked sandfishes

**** Subfamily †Judeichthyinae

:::: ''[[Apulichthys]]'' (extinct) - primitive sister group to [[Gonorynchidae|Gonorynchoids]].

***** †''[[Judeichthys]]'' Gayet 1985

:: Family '''[[Gonorynchidae]]'''

**** Subfamily Gonorynchinae

:::: ''[[Notogoneus]]'' (extinct) - from North America, Europe, Australia; some freshwater; Late Cretaceous to Oligocene; sister group to ''Gonorynchus''

***** †''[[Lecceichthys]]'' Taverne, 1998

:::: ''[[Charitosomus]]'' (extinct) - Cretaceous Gonorynchid

***** †''[[Notogoneus]]'' Cope 1885 - from North America, Europe, Australia; some freshwater; Late Cretaceous to Oligocene (=†''Anormurus'' [[Henri Marie Ducrotay de Blainville|Blainville]], 1818, †''Protocatostomus'' Whitfield 1891)

:::: ''[[Judeichthys]]'' (extinct)

***** ''[[Gonorynchus]]'' Gronow 1763 ex Scopoli 1777

:::: ''[[Ramallichthys]]'' (extinct)

** '''Suborder Chanoidei''' Berg 1937

*** Family '''[[Chanidae]]''' Günther 1868 (milkfishes)

:::: ''[[Charitopsis]]'' (extinct)

:::: ''[[Gonorynchus]]''

**** †''[[Cabindachanos]]'' Taverne et al. 2019 - [[Paleocene]]<ref>{{Cite journal |last=Taverne |first=Louis |last2=Putter |first2=Thierry DE |last3=Mees |first3=Florias |last4=Smith |first4=Thierry |date=2019-01-01 |title=Cabindachanos dartevellei gen. and sp. nov., a new chanid fish (Ostariophysi, Gonorynchiformes) from the marine Paleocene of Cabinda (Central Africa) |url=https://popups.uliege.be/1374-8505/index.php?id=6129 |journal=Geologica Belgica |language=en |doi=10.20341/gb.2018.011 |issn=1374-8505|doi-access=free }}</ref>

: Suborder '''Knerioidei'''

**** ?†''[[Coelogaster]]'' Eastman, 1905 - Eocene<ref name=":0" />

:: Family '''[[Kneriidae]]'''

**** Subfamily †[[Rubiesichthyinae]] Poyata-Ariza 1996

:::: ''[[Cromeria]]''

***** Genus †''[[Gordichthys]]'' Poyata-Ariza 1994- Early Cretaceous Chanid

:::: ''[[Grasseichthys]]''

***** Genus †''[[Nanaichthys]]'' Amaral & Brito 2012

:::: ''[[Kneria]]''

***** Genus †''[[Rubiesichthys]]'' Wenz 1984 - Early Cretaceous Chanid

:::: ''[[Parakneria]]''

**** Subfamily Chaninae

:: Family '''[[Phractolaemidae]]'''

***** Genus †''[[Aethalionopsis]]'' Gaudant 1967 - Early [[Cretaceous]]<ref>{{Citation |last=Diogo |first=Francisco José Poyato-Ariza, Terry Grande, Rui |title=Gonorynchiform Interrelationships: Historic Overview, Analysis, and Revised Systematics of the Group |date=2010 |work=Gonorynchiformes and Ostariophysan Relationships |url=https://www.taylorfrancis.com/chapters/edit/10.1201/b10194-7/gonorynchiform-interrelationships-historic-overview-analysis-revised-systematics-group-francisco-jos%C3%A9-poyato-ariza-terry-grande-rui-diogo |access-date=2024-02-06 |publisher=CRC Press |doi=10.1201/b10194-7/gonorynchiform-interrelationships-historic-overview-analysis-revised-systematics-group-francisco-jos%C3%A9-poyato-ariza-terry-grande-rui-diogo |isbn=978-0-429-06156-1}}</ref>

:::: ''[[Phractolaemus]]''

***** Genus †''[[Caeus]]'' Costa 1860 - Early Cretaceous<ref>{{Cite journal |last=Taverne |first=Louis |last2=Capasso |first2=Luigi |date=2017 |title=Osteology and relationships of Caeus ( “ Chanos ” ) leopoldi ( Teleostei , Gonorynchiformes , Chanidae ) from the marine Albian ( Early Cretaceous ) of Pietraroja ( Campania , southern Italy ) |url=https://museodistorianaturale.comune.verona.it/media/_Musei/_StoriaNaturale/_Allegati/Biblioteca/Bollettino/Bollettino%2041(2017)/Geologia/01_Taverne_Capasso_bassa.pdf |journal=Bollettino del Museo Civico di Storia Naturale di Verona |volume=41 |pages=03-20}}</ref>

***** Genus ''[[Chanos chanos|Chanos]]'' Lacépède 1803

***** Genus †''[[Dastilbe]]'' Jordan 1910 - Early Cretaceous Chanid

***** Genus †''[[Francischanos]]'' Ribeiro ''et al.'' 2022 - Early Cretaceous Chanid<ref>{{Cite journal |last=Ribeiro |first=Alexandre C. |last2=Bockmann |first2=Flávio A. |last3=Poyato-Ariza |first3=Francisco J. |date=2022-07-01 |title=Francischanos, a replacement genus for Dastilbe moraesi Silva-Santos, 1955, from the Quiricó Formation, Lower Cretaceous of the Sanfranciscana basin, Brazil (Ostariophysi: Gonorynchiformes) |url=https://www.sciencedirect.com/science/article/pii/S0195667122000763 |journal=Cretaceous Research |volume=135 |pages=105212 |doi=10.1016/j.cretres.2022.105212 |issn=0195-6671}}</ref>

***** Genus †''[[Parachanos]]'' Arambourg & Schneegans 1935 - Early Cretaceous Chanid

***** Genus †''[[Tharrhias]]'' Jordan & Branner 1908 - Early Cretaceous Chanid

***** Genus †''[[Vango (fish)|Vango]]'' Murray ''et al.'' 2023 - Late Cretaceous Chanid<ref>{{Cite journal |last=Murray |first=Alison M. |last2=Brinkman |first2=Donald B. |last3=Friedman |first3=Matt |last4=Krause |first4=David W. |date=2023-10-17 |title=A large, freshwater chanid fish (Ostariophysi: Gonorynchiformes) from the Upper Cretaceous of Madagascar |url=http://dx.doi.org/10.1080/02724634.2023.2255630 |journal=Journal of Vertebrate Paleontology |doi=10.1080/02724634.2023.2255630 |issn=0272-4634}}</ref>

** '''Suborder Knerioidei'''

*** '''Family [[Kneriidae]]''' Günther 1868

**** Genus †''[[Mahengichthys]]'' Davis, Arratia & Kaiser 2013

**** Subfamily Kneriinae (Shellears)

***** Tribe Cromeriini

****** Genus ''[[Cromeria]]'' Boulenger 1901

****** Genus ''[[Grasseichthys]]'' Géry 1964

***** Tribe Kneriini

****** Genus ''[[Kneria]]'' Steindachner 1866

****** Genus ''[[Parakneria]]'' Poll 1965

*** Family '''[[Phractolaemidae]]'''

**** Genus ''[[Phractolaemus]]'' Boulenger 1901 (Hingemouths/snake mudheads)



==Timeline of genera==

==Timeline of genera==

Line 164: Line 118:

bar:NAM16

bar:NAM16

bar:NAM17

bar:NAM17




bar:space

bar:space

Line 200: Line 155:

color:earlycretaceous bar:NAM5 from: -125 till: -112 text: [[Tharrhias]]

color:earlycretaceous bar:NAM5 from: -125 till: -112 text: [[Tharrhias]]

color:earlycretaceous bar:NAM6 from: -125 till: -93.5 text: [[Caeus]]

color:earlycretaceous bar:NAM6 from: -125 till: -93.5 text: [[Caeus]]

color:latecretaceous bar:NAM7 from: -99.6 till: -97.6 text: [[Charitopsis (fish)|Charitopsis]]

color:latecretaceous bar:NAM7 from: -99.6 till: -97.6 text: [[Charitopsis]]

color:latecretaceous bar:NAM8 from: -99.6 till: -97.6 text: [[Judeichthys]]

color:latecretaceous bar:NAM8 from: -99.6 till: -97.6 text: [[Judeichthys]]

color:latecretaceous bar:NAM9 from: -99.6 till: -97.6 text: [[Ramallichthys]]

color:latecretaceous bar:NAM9 from: -99.6 till: -97.6 text: [[Ramallichthys]]

Line 209: Line 164:

color:eocene bar:NAM14 from: -55.8 till: -33.9 text: [[Coelogaster]]

color:eocene bar:NAM14 from: -55.8 till: -33.9 text: [[Coelogaster]]

color:eocene bar:NAM15 from: -55.8 till: -23.03 text: [[Notogoneus]]

color:eocene bar:NAM15 from: -55.8 till: -23.03 text: [[Notogoneus]]

color:eocene bar:NAM16 from: -55.8 till: 0 text: [[Chanos chanos|Chanos]]

color:eocene bar:NAM16 from: -55.8 till: 0 text: [[Chanos]]

color:oligocene bar:NAM17 from: -33.9 till: -28.4 text: [[Neohalecopsis]]

color:oligocene bar:NAM17 from: -33.9 till: -28.4 text: [[Neohalecopsis]]




PlotData=

PlotData=

Line 234: Line 190:

</timeline>

</timeline>



== References ==

{{Reflist}}

* {{cite journal|last=Sepkoski |first=Jack |title=A compendium of fossil marine animal genera |journal=Bulletins of American Paleontology |volume=364 |page=560 |year=2002 |url=http://strata.ummp.lsa.umich.edu/jack/showgenera.php?taxon=611&rank=class |access-date=2011-05-17 |url-status=dead |archive-url=https://web.archive.org/web/20110723131237/http://strata.ummp.lsa.umich.edu/jack/showgenera.php?taxon=611&rank=class |archive-date=2011-07-23 }}



== References ==

{{reflist}}

* {{cite journal | last = Sepkoski | first = Jack | title = A compendium of fossil marine animal genera | journal = Bulletins of American Paleontology | volume = 364 | pages = p.560 | year = 2002 | url = http://strata.ummp.lsa.umich.edu/jack/showgenera.php?taxon=611&rank=class | accessdate = 2011-05-17 }}

{{Actinopterygii}}

{{Actinopterygii}}

{{Taxonbar|from=Q755012}}

{{Authority control}}



[[Category:Gonorynchiformes| ]]

[[Category:Gonorynchiformes| ]]

[[Category:Extant Berriasian first appearances]]

[[Category:Ostariophysi]]


[[Category:Ray-finned fish orders]]

{{Actinopterygii-stub}}

[[Category:Taxa named by Lev Berg]]


[[af:Gonorynchiformes]]

[[az:Burnukonuskimilər]]

[[ca:Gonorinquiforme]]

[[de:Sandfischartige]]

[[es:Gonorynchiformes]]

[[fr:Gonorynchiformes]]

[[gl:Gonorrinchiformes]]

[[ko:압치목]]

[[lt:Arkliagalvės]]

[[hu:Gonorynchiformes]]

[[nl:Zandvisachtigen]]

[[ja:ネズミギス目]]

[[oc:Gonorynchiformes]]

[[pl:Piaskolcokształtne]]

[[pt:Gonorynchiformes]]

[[ru:Гоноринхообразные]]

[[fi:Maitokalat]]

[[th:อันดับปลานวลจันทร์ทะเล]]

[[tr:Gonorynchiformes]]

[[zh:鼠鱚目]]

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