Contents

Ursinae

Ursinae Temporal range: Barstovian–Recent PreꞒ Ꞓ O S D C P T J K Pg N
AEurasian brown bear (Ursus arctos arctos)
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Mammalia
Order:	Carnivora
Family:	Ursidae
Subfamily:	Ursinae Fischer de Waldheim, 1817
Genera
See text.

Ursinae is a subfamily of Ursidae (bears) named by Swainson (1835). It was assigned to Ursidae by Bjork (1970), Hunt (1998), and Jin et al. (2007).^[1][2][3]

The genera Melursus and Helarctos are sometimes also included in Ursus. The Asiatic black bear and the polar bear used to be placed in their own genera, Selenarctos and Thalarctos; these are now placed at subgenus rank.

• Subfamily Ursinae Fischer de Waldheim, 1817
  • †Aurorarctos Jiangzuo & Flynn, 2020
    • †Aurorarctos tirawa Jiangzuo & Flynn, 2020
  • Helarctos Horsfield, 1825
    • Helarctos malayanus (Raffles, 1821) – sun bear
    • †Helarctos sinomalayanus (Thenius, 1947)
  • Melursus Meyer, 1793
    • Melursus ursinus (Shaw, 1791) – sloth bear
    • †Melursus theobaldi (Lydekker, 1884)
  • †Protarctos Kretzoi, 1945
    • †Protarctos abstrusus (Bjork, 1970)
    • †Protarctos boeckhi (Schlosser, 1899)
    • †Protarctos ruscinensis (Depéret, 1890)
    • †Protarctos yinanensis (Li, 1993)
  • Ursus Linnaeus, 1758
    • Ursus americanus (Pallas, 1780) – American black bear
    • Ursus arctos Linnaeus, 1758 – brown bear
    • †Ursus deningeri Richenau, 1904
    • †Ursus dolinensis (Garcia & Arsuaga, 2001)
    • †Ursus etruscus Cuvier, 1823
    • †Ursus ingressus Rabeder, Hofreiter, Nagel & Withalm 2004
    • †Ursus kudarensis Baryshnikov, 1985
    • Ursus maritimus Phipps, 1774 – polar bear
    • †Ursus minimus (Devèze & Bouillet, 1827)
    • †Ursus pyrenaicus (Depéret, 1892)
    • †Ursus rossicus Borissiak, 1930
    • †Ursus sackdillingensis Heller, 1955
    • †Ursus savini (Andrews, 1922)
    • †Ursus spelaeus Rosenmüller, 1794 – cave bear
    • Ursus thibetanus (Cuvier, 1823) – Asiatic black bear
    • †Ursus vitabilis?Gidley, 1913

A number of hybrids have been bred between American black, brown, and polar bears (see Ursid hybrids).

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